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Razowski, J.
Tortricidae from South Africa, 4: Neopotamia - group of Olethreutini (Lepidoptera:
Tortricidae)
SHILAP Revista de Lepidopterología, Vol. 36, Núm. 141, marzo-sin mes, 2008, pp. 57-
68
Sociedad Hispano-Luso-Americana de Lepidopterología
España

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SHILAP Revista de Lepidopterología
ISSN (Versión impresa): 0300-5267
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Sociedad Hispano-Luso-Americana de
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SHILAP Revta. lepid., 36 (141), marzo 2008: 57-68 CODEN: SRLPEF ISSN:0300-5267

Tortricidae from South Africa, 4:
Neopotamia - group of Olethreutini
(Lepidoptera: Tortricidae)
J. Razowski

Abstract

Three genera (Promodra Razowski, gen. n., Rufeccopsis Razowski, gen. n., Zellereccopsis Razowski, gen.
n.) and 6 species (Eccopsis ofcolacona Razowski, sp. n., Zellereccopsis caffreana Razowski, sp. n., Promodra
nigrata Razowski, sp. n., Paraeccopsis windhoeca Razowski, sp. n., Afroploce mabalinguae Razowski, sp. n.,
Rufeccopsis brunneograpta Razowski, sp. n.) are described as new; 6 new combinations are proposed; new
distribution data are provided for all known species.
KEY WORDS: Lepidoptera, Tortricidae, Olethreutini, Neopotamia, new taxa, Afrotropical.

Tortricidae del sur de África, 4: grupo - Neopotamia de Olethreutini
(Lepidoptera: Tortricidae)

Resumen

Se describen como nuevas tres géneros (Promodra Razowski, gen. n., Rufeccopsis Razowski, gen. n.,
Zellereccopsis Razowski, gen. n.) y 6 especies (Eccopsis ofcolacona Razowski, sp. n., Zellereccopsis caffreana
Razowski, sp. n., Promodra nigrata Razowski, sp. n., Paraeccopsis windhoeca Razowski, sp. n., Afroploce
mabalinguae Razowski, sp. n., Rufeccopsis brunneograpta Razowski, sp. n.); se proponen 6 nuevas
combinaciones; se proveen datos de distribución para todas las especies conocidas.
PALABRAS CLAVE: Lepidoptera, Tortricidae, Olethreutini, Neopotamia, nuevas taxa, Afrotropical.

Introduction

DIAKONOFF (1973) grouped 11 genera of Olethreutini in his subtribe Neopotamiae then
(DIAKONOFF, 1983) he added his Madagascan Xenopotamia. RAZOWSKI (1989) included
Neopotamiae in Olethreutini and HORAK (2004) treated them as a species group of that tribe and
provided a comprehensive redefinition. AARVIK (2004) discussed the Afrotropical Neopotamiae
and placed in them Eccopsis Zeller, 1852, Cosmorryncha Meyrick, 1913, Metendothenia Diakonoff,
1973, Megalota Diakonoff, 1966, Basigonia Diakonoff, 1983 and the following six new genera:
Afrocostosa, Neorrhyncha, Geita, Paraeccopsis, Afrothreutes and Afroploce and 14 new species.
Altogether he listed as much as 28 Afrotropical species of this group. In this paper 2 known
Afrotropical genera are discussed and three new genera are described. Now, Neopotamia-group of
genera is represented in the tropical Africa by 14 genera and 45 species and one can suppose that it
is the largest group of Olethreutini in this region. The geographic distribution of this group is
pantropical; the most abundant it is in the Oriental and Afrotropical regions, innumerous taxa (4
genera and 6 species) are Australian (HORAK, 2004) and only two genera (Eccopsis, Megalota)

57
J. RAZOWSKI

and about 10 species are discovered in the Neotropical region; in Palaearctic the Neopotamia -
group is represented by Metendothenia and a single species of Eccopsis (in Saudi Arabia) only.
Abbreviations. GS - genitalia slide; ISEZ - Institute of Systematics and Evolution of Animals
PAS, Kraków; RSA - Republic South Africa.

SYSTEMATICS

Eccopsis Zeller, 1852
This genus was redescribed by DIAKONOFF (1981) and AARVIK (2004) and compared with
Cosmorryncha Meyrick, 1913. These two genera and Metendothenia Diakonoff, 1973 are very
close to one the other. DIAKONOFF mentions a few differing characters but they are not
convincing. AARVIK (2004) placed two externally very similar species in two genera,
Cosmorrhyncha and his Neorrhyncha. Very similar facies is now found in several Neotropical
species and some of them certainly belong to Eccopsis. It thus seems possible that Cosmorryncha
and Eccopsis are synonymous.

Eccopsis ptilonota (Meyrick, 1921)
Material examined. One male from Pretoria (1- XII-1911, Capt. Paget).
Remarks. Holotype, male, was described and illustrated by RAZOWSKI & KRÜGER (2007).

Eccopsis praecedens Walsingham, 1897
Material examined. One male from Nelspruit (12-III-1967, Potgieter & Goode).
Remarks. E. praecedens was described from French Congo. AARVIK (2004) recorded it from
Angola, Zaire, Kenya, Nigeria, Tanzania, Reunion and South Africa (Pinetown, Natal and
Wilderness, Cape Province). Its synonymy, E. chromatica Diakonoff, 1983 is from Cape Verde
Islands.

Eccopsis wahlbergiana Zeller, 1852
Material examined. One female from Silverton (VIII-1961); Knysna, Cape Province (10-14-I-
1955, A. J. T. Janse); Storms River Pass (8-III-1954, L. Vári); South Rhodesia (Lundi, 13-16-I-
1954, Vári & van Son).
Remarks. A variable species. Widely distributed: from Cameroon and Cabo Verde Islands to
Madagascar and Saudi Arabia. AARVIK (2004) records a few specimens from South Africa (Natal
and Cape Province). This species was described from South Africa, its synonymy, Eccopsis
fluctuatana Walsingham, 1881 is from Natal, same country.

Eccopsis affluens (Meyrick, 1921)
Material examined. One specimen without locality data.
Remarks. Male redescribed by RAZOWSKI & KRÜGER (2007). Female genitalia similar to
wahlbergiana and E. ochrana Aarvik, 2004 from Tanzania but affluens with long postostial part of
sterigma followed by small median sclerite. This species was described from Mozambique
(Portuguese East Africa).

Eccopsis ofcolacona Razowski sp. n. (Fig. 1)

Holotype female: “Malta Forest, Tvl. nr. Ofcolaco, 19- IX-1960, van Son & L. Vári”; GS 267
RSA. Deposited in the Transvaal Museum, Pretoria.
Description. Wing span 18 mm. Head and thorax yellowish brown. Forewing as in praecedens
and wahlbergiana. Ground colour cream ochreous; suffusions more brown; dorsum suffused with
brown.
Markings diffuse, consisting of ochreous brown basal suffusion, median fascia interrupted

58 SHILAP Revta. lepid., 36 (141), marzo 2008
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subcostally, included in brown suffusion at dorsum; subterminal fascia very slender, brownish;
costal spots brownish. Cilia concolorous with ground colour, brownish to 2/3 of termen. Hindwing
pale brownish, ochreous along outer edge; anal area not differentiated; cilia brownish.
Male not known.
Female genitalia (Fig. 12). Papilla analis broadest medially; apophyses posteriores long; plates
of sterigma moderate; antrum long, in distal 2/3 uniformly broad, sclerotized, in proximal portion
weakly so; ductus bursae slender; signum with two distinct blades.
Diagnosis. Facies similar to ochreana and E. tucki Aarvik, 2004 from Kenya but ofcolacona
with forewing termen distinctly concave beyond apex (approximately at vein M2), brownish
hindwing and uniformly broad distal half of sclerite of antrum.
Etymology. The name refers to the type locality.

Eccopsis incultana (Walker, 1863)
Material examined. Two females from Blyde River Nature Reserve (24-I - 1-XII-1975,
Potgiter & Scoble) and one male from Satara (29-IV - 2-V-1964, Potgieter & Goode).
Remarks. Widely distributed from Zaire to Tanzania; from South Africa recorded by AARVIK
(2004) from Natal and Zimbabwe. Described from Mauritius; its synonymies are: Argyroploce
trixiphias Meyrick, 1939 from Belgian Congo (Zaire) and E. undosa Diakonoff, 1981 from
Madagascar.

Zellereccopsis Razowski, gen. n.

Type-species: Zellereccopsis caffreana Razowski, sp. n.
Description. Venation. In forewing chorda fully developed, terminating beneath R5; M-stem
terminating before M2; R5 to beneath apex; CuA2 opposite mid-distance between R1-R2. In
hindwing Rr - M1 approaching to one another to middle; M3-CuA1 connate; M2 well separate from
base of the latter.
Male genitalia. Uncus long, broadest medially; socius broad to middle, tapering termined,
slightly curved apically, without hairs dorsally and basally; gnathos ill-defined; valvae rather
symmetric with indistinct neck; process of posterior edge of basal cavity strongly reduced; cucullus
elongate with indistinct ventroproximal angle from which a row of rather strong setae extends;
aedeagus slender, bent, with dorso-subterminal thorn; one slender, fairly long cornutus in vesica.
Female not known.
Distribution. South Africa. A monotypic genus.
Diagnosis. The new genus is related to Eccopsis Zeller, 1852 as shapes of the tegumen and
valva and the situation of socii show. Zellereccopsis differs from Eccopsis in constricted base of
uncus, sparsely hairy, pointed termination of uncus; slender terminal parts of socius and reduction
of subdorsal process of distal edge of basal cavity of valva.
Etymology. The generic name refers to the name Eccopsis and the name of its author, E. P.
Zeller.

Zellereccopsis caffreana Razowski, sp. n. (Fig. 2)

Holotype male: “Skukuza K. N. P. Survey, 13-13- II-1963, L. Vári”; GS 295 RSA. Deposited
in the Transvaal Museum, Pretoria.
Description. Wing span 16.5 mm. Head and thorax pale ochreous cream. Forewing expanding
termined; termen not oblique, straight. Ground colour ochreous cream with slight pinkish
admixture; costal divisions, lines and strigulae brownish. Basal blotch weakly developed marked
with some spots strongest dorsoposteriorly. Median fascia discontinuous medially, followed by
weak grey-brown convex fascia; subterminal fascia slender. Cilia concolorous with ground colour

SHILAP Revta. lepid., 36 (141), marzo 2008 59
J. RAZOWSKI

with browner scales. Hindwing brownish in distal half mixed rust, with anal portion convex. Cilia
cream mixed with rust mainly in apical part.
Male genitalia (Fig. 7) as described for the genus.
Diagnosis. The only species of the genus (see genus description). Externally similar to E.
praecedens.
Etymology. The specific name refers to Africa, Terra Caffrorum.

Cosmorryncha microcosma Aarvik, 2004
Material examined. One female from Graskop (20-21-II-1962, Vári & Leleup); 3 specimens:
Mt. Selinda (9-17-IV-1956, L. Vári); Fourteen Streams, Barberton District (13-15-III-1967,
Potgieter & Goode).
Remarks. This species was described from Zaire, Belgian Congo (holotype), Kenya, São
Thome and Uganda. The present male differs from those in the figure by Aarvik in broader,
rounded terminally socii and slightly longer, slenderer antrum.

Metendothenia balanacma (Meyrick, 1914)
Material examined. One female from Gobabeb, Game Reserve No. 3, (20-29-V-1965, J. H.
Potgieter); a pair from Abachans (V-1944, G. Hobohm); both localities in Swaziland.
Remarks. RAZOWSKI & KRÜGER (2007) illustrated the holotype of both balanacma (from
East Africa) and its synonym Argyroploce anaclina Meyrick, 1921 (from Rhodesia). AARVIK
(2004) redescribed this species and provided the data on its distribution: Kenya, Malawi,
Mozambique, Namibia, Tanzania and Zimbabwe.

Geita micrograpta (Meyrick, 1921), comb. n.
Remarks. RAZOWSKI & KRÜGER (2007) redescribed and illustrated the male holotype of
micrograpta (from East Africa) in its original genus, Argyroploce. AARVIK (2004) established a
new monobasic genus Geita for his G. bioernstadi from Tanzania and Congo (Zaire). G. bioernstadi
differs from micrograpta in large potsangular bunch of spines of right sacculus and larger spine of
dorsoposterior edge of basal cavity of valva. The presence of dorsal thorns of postmedian part of
aedeagus in micrograpta are of lesser importance.

Megalota sponditis (Meyrick, 1918)
Material examined. One male from Kwa Zulu-Natal (Ngome, 18-22. I. 1993, M. Krüger( and
three females from Umdoni Park (6 -12-I-1976, D. M. Kroon); 8 miles North of Livingstone (25-
28-V-1954, A. J. T. Janse), and Umzinto National Park (24-XII-1930, A. J. T. Janse).
Male genitalia as in M. lobotona (cf. below).
Female genitalia (Fig. 13). Posterior part of sterigma with pair of small median lobes; antrum
large, partially well sclerotized, tapering proximed; signum with three small processes.
Remarks. This species was described from Cape Colony; the holotype male (with abdomen
missing) was illustrated by RAZOWSKI & KRÜGER (2007). It is very close to M. purpurana
Aarvik, 2004 from Kenya known also from females but in spondytis antrum is more slender.

Megalota lobotona (Meyrick, 1921)
Material examined. Two females from Pretoria (21-IX-1911, A. T. J. Janse), and Swaziland
(Abachans, II-1944, G. Hobohm).
Female genitalia (Fig. 14). Sterigma weak with small but well preserved median prominences;
antrum large, rather uniformly broad except for small proximal portion; signum with small lobes.
Remarks. RAZOWSKI & KRÜGER (2007) illustrated male holotype.
This species was illustrated (adult and male) by RAZOWSKI & KRÜGER 2007). Facies of the
lobotona and sponditis are different whilst the genitalia very close. This was observed by AARVIK
(2004) in the case of other four species (M. rhopalitis Meyrick, 1920, and his M. namibiana,

60 SHILAP Revta. lepid., 36 (141), marzo 2008
TORTRICIDAE FROM SOUTH AFRICA, 4: NEOPOTAMIA - GROUP OF OLETHREUTINI

purpurana and archana of 2004) which differ in their aedeagi. In lobotona and namibiana aedeagus
terminates in a dorsal thorn and in sponditis and purpurana in a ventrosubterminal thorn. Female of
lobotona differs from sponditis in its broader antrum.

Promodra Razowski, gen. n.

Type-species: Argyroploce prodroma Meyrick, 1913
Description. Venation. In forewing all veins separate, median stem absent, chorda from 1/4 of
R1-R2; CuA2 opposite 3/4 distance between bases of R1-R2; in hindwing M3-CuA1 connate,
remaining veins separate.
Description. Male genitalia (figured by RAZOWSKI & KRÜGER, 2007). Basal and terminal
parts of uncus slender; row of spines along posteromedian part of ventral surface of uncus; valvae
symmetric with broad neck and subsquare cucullus; two groups of bristles, one above angle of
sacculus, the other (larger spines) before cucullus, subdorsally; aedeagus well sclerotized, slender,
broadest subterminally.
Female genitalia (description based on prodroma). Papillae anales small, slender; apophyses
slender; sterigma membranous; ostium bursae broad, antrum broad, distinctly sclerotized around
ostium; remaining parts of ductus bursae slender, elongate sclerite in a loop well before middle;
signum a half-moon-shaped sclerite.
Distribution. Known from South Africa.
Diagnosis. Close to Paraeccopsis as the shape of valvae and its mediosubdorsal group of
spines show; Promodra is distinct by large basally constricted and terminally pointed uncus with
ventromedian arrangement of setae and short, transverse pocket like signum.
Etymology. The generic name is an anagram of the name of type-species.

Promodra prodroma (Meyrick, 1913)
Material examined. “Mariti Forest, 6-7- III-1974, L. & G. Vári”; Barbarton (25- I-1911, A. T.
J. Janse), and paratypes.
Variation. Specimen from Mariti Forest with diffuse, blackish median fascia, brownish,
strigulated with blackish brown proximal area and cream ferruginous ground colour of terminal
area.
Remarks. Male genitalia described by RAZOWSKI & KRÜGER (2007), for description of
female genitalia (Fig. 15) see characteristics of the genus. Described from South Africa.

Promodra nigrata Razowski, sp. n. (Fig. 3)

Holotype female: “Harkerville Forest Reserve, 1-5-III-1972, L. & G. Vári”; GS 334 RSA.
Deposited in the Transvaal Museum, Pretoria.
Description. Wing span ca 14 mm; head and thorax pale greyish brown, vertex more cream,
base of tegula brown. Forewing slightly expanding terminally; termen weakly oblique, straight.
Ground colour brownish cream; suffusions, strigulae, and dots brownish; basal blotch grey brown
with darker marks chiefly at costa posteriorly; median fascia blackish brown in costal half of wing,
otherwise paler, with brown spots followed by greyer postmedian area; subterminal fascia reduced
to oval blotch accompanied by brown costal spots. Cilia brown-grey, glossy. Hindwing pale greyish
brown with cilia similar.
Male not known.
Female genitalia (Fig. 16). Papillae anales moderate; sterigma ill-defined; antrum large, bent,
distinctly sclerotized, uniformly so towards ostium bursae; proximal half of ductus bursae slender,
without any sclerite; signum an oval pocket with distinct half-moon -shaped ventroproximal
portion.

SHILAP Revta. lepid., 36 (141), marzo 2008 61
J. RAZOWSKI

Diagnosis. Facies reminiscent of prodroma but nigrata with termen of forewing more oblique
and antrum long, distinctly sclerotized in distal half, not tapering proximally.
Etymology. The specific name refers to maculation of forewing; Latin: nigrata - mixed black.

Paraeccopsis Aarvik, 2004
Monotypic Paraeccopsis Aarvik, 2004 was described to comprise Argyroploce insellata
(Meyrick, 1920). Six further species are included in Paraeccopsis in this paper. The systematic
status of four Paraeccopsis nucleata (Meyrick, 1913), comb. n., Paraeccopsis exhilarata (Meyrick,
1918), comb. n., Paraeccopsis acroplecta (Meyrick, 1921), comb. n., Paraeccopsis eoplecta
(Meyrick, 1925), comb. n., of them is somewhat unclear. Both the facies and female genitalia of
these species are almost identical and their males remain unknown. The male genitalia of three
closely related species, P. phoeniodes (Meyrick, 1921), comb. n., P. acroplecta and P. insellata are
easily distinguished from one the other. The female of the latter is extremely similar to P. nucleata
and the two following species. It thus is supposed that these species are synonymous. P. phoeniodes
and P. insellata are distinct externally. Unfortunately P. insellata and two its synonymies (=
Polychrosis inflicta Meyrick, 1920 from India and Argyroploce atricapsis Meyrick, 1930 from
Nigeria) sensu AARVIK 2004 are known from females only.

Paraeccopsis insellata (Meyrick, 1920)
Material examined. Mozambique (Maronga Forest 20º 03’ S, 33º 09’ E, 6-11-IX-1972, R. H.
Jones).
Remarks. Known from Botswana, Gambia, Kenya, Nigeria, Tanzania; recorded also from the
Oriental Region: India. Synonymies: Polychrosis inflicta Meyrick, 1920 (Bombay, India) and
Argyroploce atricapitis Meyrick 1930 (Nigeria (AARVIK 2004)).

Paraeccopsis acroplecta (Meyrick, 1921)
Material examined. A pair labelled: Zululand (I-1948, J. C. Faure) and Gamtoos River (28-
XII-1951, C. & C. Dickson).
Description. Female genitalia (Fig. 17). Distal, sclerotized part of ductus bursae as long as its
membranous proximal portion; cup-shaped part of sterigma; medioposterior slender part of sterigma
situated between large lateral plates; signum with three equally sized processes.
Remarks. Male holotype (from Rhodesia) was illustrated and discussed by RAZOWSKI &
KRÜGER (2007). The identification of specimens was based on a comparison with the type
material. Compare the discussion for Paraeccopsis. Female was unknown to this date.

Paraeccopsis phoeniodes (Meyrick, 1921)
Material examined. Illwo Beak (25-XI-1986, L. Vári, on Acacia). Type redescribed by
RAZOWSKI & KRÜGER 2007) under the original generic name.

Paraeccopsis windhoeca Razowski, sp. n. (Fig. 4)

Holotype male: “Windhoek, 26-29-I-1951, R. G. Stery”; GS 186 RSA. Deposited in the
Transvaal Museum, Pretoria.
Description. Wing span 15 mm. Front cream slightly tinged with brownish, thorax darker.
Forewing weakly expanding posteriorly; costa not convex; termen fairly oblique, almost straight.
Ground colour cream brownish suffused and strigulated with brownish; dorsal patch suffused with
ochreous. Basal blotch ill-defined, brownish with brown posterior spots; costal third of median
fascia brownish, remaining parts brownish ochreous; subterminal fascia concolorous, preserved at
dorsum; brown spot at apex of wing; divisions pale brownish. Cilia brownish cream scaled with
brownish. Hindwing brownish cream, whiter basad; cilia dirty white.
Male genitalia (Fig. 8). Uncus broad basally, constricted postmedially, somewhat expanding

62 SHILAP Revta. lepid., 36 (141), marzo 2008
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and spined terminally; socius slender; neck of valva developed; cucullus oval with weak
ventroproximal lobe; large dorsal lobe of posterior edge of basal cavity armed with numerous thin,
long spines; terminal portion of aedeagus slightly curved upwards.
Diagnosis. Facies somewhat similar to insellana but forewing markings ill-defined, hindwing
pale brownish cream, ventral termination of aedeagus pointed directed upwards, and lobe of
posterior edge of basal cavity of valva long bristeld.
Etymology. The name refers to the type locality.

Afroploce karsholti Aarvik, 2004
Material examined. One pair from Mt. Selinda (9-17- IV-1956, L. Vári) and one male from
Xiluvo, Moc.[ambique], VIIa Machado District (9-10- III-1964, Vári & Van Son).
Remarks. This species was described from Tanzania (holotype), Congo (Zaire), Ghana, Kenya,
and Malavi. The present specimen with the spines of the angular group of sacculus longer than
those in the original illustration by AARVIK (2004). New for South Africa.

Afroploce mabalingwae Razowski, sp. n. (Fig. 5)

Holotype male: “S. Afr.: N Province Mabalingwe 25 km W Warmbaths SE 24 28 Cd; 21-III-
1996 B. Dombrowsky, ligh trap”; GS 311 RSA. Deposited in the Transvaal Museum, Pretoria.
Paratypes 5 1 1, 3 0 0 labelled: “South Africa, Royal Natal National Park, Tendele camp,
1600 m, 6-13-XII-2004 Marek Kopec leg. (1 1; G. S. 13766) and similar label, dates 23-X, 25-X,
27-X-2003, at light, leg. L. Przybylowicz; one male from Drakensberg Park, Cathedral Peak,
Didima, 1400 m, 26-XI - 2-XII-2004, Marek Kopec leg. Paratypes in the collection of ISEZ.
Description. Wing span 18 mm. Head brown; posterior half of labial palpus black; thorax
brown-grey, tegula brown proximally. Forewing somewhat expanding posteriorly; costa weakly
convex; apex rounded; termen weakly oblique, hardly convex. Ground colour white, slightly tinged
with pinkish subterminally, with greyish near costa; some refractive bluish spots present. Suffusions
and spots brownish and blackish. Markings indistinct, blackish grey with some brownwer parts.
Cilia grey, blackish terminally with some similar divisions. Hindwing pale brownish grey; cilia
cream.
Male genitalia (Figs 9, 10). Uncus short; socii moderately broad; valva broad to middle with
short, broad hairy lobe of posterior edge of basal cavity; sacculus gently concave near middle,
rounded caudally, with long group of dense setae above ventral edge and two stout spines at base of
cucullus; cucullus moderately broad somewhat narrowing postmedially; aedeagus fairly broad,
rather short; group of 6 short cornuti in vesica.
Female genitalia (Fig. 17). Sclerite surrounding ostium bursae large, postostial sterigma fairly
long with slender lateroterminal lobes; antrum in major part sclerotized with distinct mediolateral
diverticle; signum with single proximal lobe.
Diagnosis. Closely related to A. turiana Aarvik, 2004 from Kenya but mabalinguae with two
short spines above end of sacculus and large group of setae above its distal fourth.
Etymology. The specific name refers to the type locality.

Rufeccopsis Razowski, gen. n.

Type-species: Eucosma rufescens Meyrick, 1914
Description. Venation. In forewing R5 to beneath apex; chorda from 1/4 id R1-R2 to beyond
R5, weak in its 2/3; CuA2 opposite mid-distance between R1-R2. In hindwing Rr M1 approached to
middle; M3 - CuA1 stalked to 1/3; M2 far from the latter.
Male genitalia (described and figured by RAZOWSKI & KRÜGER, 2007). Tegumen
proportionally small; socii long, well sclerotized rods; tuba analis in major part well sclerotized;
right valva slender, sparsely hairy; sacculus long, deeply concave postmedially, with free

SHILAP Revta. lepid., 36 (141), marzo 2008 63
J. RAZOWSKI

termination; left valva very broad, oval with group of setae at the end of sacculus; this latter simple,
convex; both valvae with large, capitate processes of distal edge of basal cavity; aedeagus long; two
rows of short spines on vesica.
Female genitalia of rufescens (Fig. 19). Apophyses slender, moderately long; sterigma
membranous except for slender, asymmetric anteostial part; antrum membranous, asymmetric;
bursa copulatrix membranous, without signum; ductus seminalis originating in proximal portion of
ductus bursae.
Diagnosis. Rufeccopsis is the most specialized genus of the Neopotamia -group of genera
characterized by very strong asymmetry of valvae. In shape of socii Rufeccopsis resembles
Afrocostosa Aarvik, 2004 in which, however, the valvae are symmetric and aedeagus short.
Asymmetry of valvae is well expressed in some Metendothenia Diakonoff, 1973 (e.g. in M.
balanacma (Meyrick, 1914) especially in the distribution of the vestiture elements. The processes
of posterior edge of basal cavity of rufescens are very large.
Etymology. The generic name refers to the genus name Eccopsis and the name of the type-
species and Latin: rufus - rust.
Remarks. Two species known, the type species and R. brunneograpta Razowski, sp. n., both
Afrotropical.

Rufeccopsis rufescens (Meyrick, 1914), comb. n.
Material examined. The holotype female (not illustrated but discussed by RAZOWSKI &
KRÜGER (2007) genitalia on slide 338 RSA); Rhodesia (Lundi, 13-16- III-1964, Vári & van Son);
Hope, Fnln, Rhodesia (17-I-1918, A. J. T. Janse); Three Sisters (12-III-1911, A. J. T. Janse), and
two specimens with incomplete labels (P. P. Rust, 23-XII-1925, A. J. T. Janse).
Description. Female genitalia (Fig. 19) described with the genus.

Rufeccopsis brunneograpta Razowski, sp. n. (Fig. 6)

Holotype male: “Victoria Falls, Big Tree, 6-V-1954, A. J. T. Janse”; GS 176 RSA. Deposited
in the Transvaal Museum, Pretoria.
Description. Wing span 19.5 mm. Head and thorax cream brownish. Forewing weakly
expanding terminally; termen weakly concave beneath apex. Ground colour cream brownish with
slight reddish admixture creamer in basal portion subcostally; costal strigulae cream divided rust;
apex and costal third of termen finely edged with rust. Marking dark brown: weak postbasal lines in
dorsal half of wing and subterminally; dorsal blotch consisting of some parallel elements. Cilia
paler than ground colour, creamer at tornus. Hindwing yellowish cream, darker apically; cilia
concolorous with middle of wing.
Male genitalia (Fig. 11). Arms of uncus somewhat asymmetric, left shorter than the right; right
valva slender, tapering terminated in distal half; basal third of sacculus convex, postmedian edge
broadly concave, without a subterminal lobe, free termination broad, short, directed caudally; left
valva very broad, oval; process of posterior edge of basal cavity delicate; two asymmetric rows of
short thorn like cornuti present.
Diagnosis. Closely related to rufescens but brunneograpta with longer forewing and dark
brown dorsal parts of markings, weak process of posterior edge of basal cavity of valva and smaller,
directed terminally end part of right sacculus.
Etymology. The specific name refers to the colouration of forewing consisting some brown
elements; Latin: brunneus - brown and Greek: graptos - painted.

Acknowledgements

The author thanks Dr Martin Krüger, Lepidoptera Department, the Transvaal Museum, Pretoria
who kindly provided the material for the present study, Mr Witold Zajda, Kraków for dissection of

64 SHILAP Revta. lepid., 36 (141), marzo 2008
TORTRICIDAE FROM SOUTH AFRICA, 4: NEOPOTAMIA - GROUP OF OLETHREUTINI

specimens and to Mr Marek Kopec, Kraków for the photographs and digital arrangement of
illustrations.
BIBLIOGRAPHY

AARVIK, L., 2004.– Revision of the subtribe Neopotamiae (Lepidoptera: Tortricidae) in Africa.– Norw. J.
Entomol., 51: 71-122.
DIAKONOFF, A., 1973.– The South Asiatic Olethreutini (Lepidoptera, Tortricidae).– Zool. Monogr. Rijksmus.
Natuurl. Hist. 1: 1-700, 15 pls.
DIAKONOFF, A., 1981.– Tortricidae from Madagascar. Part 2. Olethreutinae, 1.– Annls Soc. ent. Fr. (N. S.)., 17(1):
7-32.
DIAKONOFF, A., 1983.– Tortricidae from Madagascar part 2. Olethreutinae, 2.– Annls Soc. ent. Fr., (N. S.)., 19:
291-310.
HORAK, M., 2004.– Olethreutinae moths of Australia (Lepidoptera: Tortricidae).– Monogr. Australian Lepid., 10:
1-522.
RAZOWSKI, J., 1989.– The genera of Tortricidae (Lepidoptera). Part II: Palaearctic Olethreutinae.– Acta zool.
cracov., 32(7): 107-328.
RAZOWSKI, J. & KRÜGER M., 2007.– An illustrated catalogue of the type specimens of Tortricidae in the
Transvaal Museum, Pretoria (Lepidoptera: Tortricidae).– SHILAP Revta. lepid., 35(138): 103-179.

J. R.
Institute of Systematics and Evolution of Animals
Polish Academy of Sciences
Slawkowska, 17
PL-31-016 Kraków
POLONIA / POLAND
E-mail: Razowski@isez.pan.krakow.pl

(Recibido para publicación/ Received for publication 11-VII-2007)
(Revisado y aceptado/ Revised and accepted 20-VIII-2007)

SHILAP Revta. lepid., 36 (141), marzo 2008 65
J. RAZOWSKI

1 2

3 4

5 6

Figs. 1-6.– Adults. 1. Eccopsis oflacolala Razowski, sp. n., holotype; 2. Zellerecopsis caffreana Razowski, sp.
n., holotype; 3. Promodra nigrata Razowski, sp. n., holotype; 4. Pareccopsis windhoeca Razowski, sp. n.,
holotype; 5. Afroploce mabalinguae Razowski, sp. n., holotype; 6. Rufeccopsis brunneograpta Razowski, sp.
n., holotype.

66 SHILAP Revta. lepid., 36 (141), marzo 2008
TORTRICIDAE FROM SOUTH AFRICA, 4: NEOPOTAMIA - GROUP OF OLETHREUTINI

7
9

8
12

11
Figs 7.– 11. Male and female genitalia: 7. Zellerecopsis caffreana Razowski, sp. n., holotype; 8. Pareccopsis
windhoeca Razowski, sp. n., holotype; 9, 10. Afroploce mabalinguae Razowski, sp. n., holotype; 11.
Rufeccopsis brunneograpta Razowski, sp. n., holotype; 12. Eccopsis oflacolala Razowski, sp. n., holotype; 13.
Megalota spondylis (Meyrick), Umzito.

SHILAP Revta. lepid., 36 (141), marzo 2008 67
J. RAZOWSKI

Figs 14.– 18. Female genitalia: 14. Megalota lobotona (Meyrick), Pretoria; 15. Promodra prodroma (Meyrick),
Barberton; 16. Promodra nigrata Razowski, sp. n., holotype; 17. Afroploce mabalinguae Razowski, sp. n.,
paratype, GS 13766; 18. Megalota acroplecta (Meyrick), Zululand. 19. Rufeccopsis rufescens (Meyrick),
holotype.

68 SHILAP Revta. lepid., 36 (141), marzo 2008