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Biology Chemistry Frontier

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Biology - Chemistry Frontier. Part 1. Roger Adams Award Lecture Samuel Danishefsky, Memorial Sloan-Kettering Cancer Center Biology - Chemistry Frontier. Part 1. Roger Adams Award Lecture Samuel Danishefsky, Memorial Sloan-Kettering Cancer Center Colleagues Multivalent Antitumor Vaccine Jennifer Allen Stacy Keding Qian Wan Erythropoietin / Peptide Ligation Gong Chen Jiehao Chen Zihao Hua Cindy Kan Zhongping Tan John Trzupek Qian Wan David Warren Bin Wu Yu Yuan Grandisine A David Maloney Migrastatin Christoph Gaul Jon Njardarson Lucy Perez Isomigrastatin Isaac Krauss Mihir Mandal Epothilones Alexey Rivkin Young Shin Cho Fumihiko Yoshimura Cycloproparadicicol Xudong Geng Kana Yamamoto Zhi-Quang Yang Colleagues Multivalent Antitumor Vaccine Jennifer Allen Stacy Keding Qian Wan Erythropoietin / Peptide Ligation Gong Chen Jiehao Chen Zihao Hua Cindy Kan Zhongping Tan John Trzupek Qian Wan David Warren Bin Wu Yu Yuan Grandisine A David Maloney Migrastatin Christoph Gaul Jon Njardarson Lucy Perez Isomigrastatin Isaac Krauss Mihir Mandal Epothilones Alexey Rivkin Young Shin Cho Fumihiko Yoshimura Cycloproparadicicol Xudong Geng Kana Yamamoto Zhi-Quang Yang Acknowledgement I dedicate this Roger Adams Award Lecture to Sarah Danishefsky for a lifetime of creative support, encouragement and leadership. Acknowledgement I dedicate this Roger Adams Award Lecture to Sarah Danishefsky for a lifetime of creative support, encouragement and leadership. Biology Intellectualization at the level of function Mechanisms: Identification of all components required to attain and maintain biological function → reconstitution! The power of bioreplicative synthesis Chemistry Intellectualization at the level of structure Mechanisms: Pathways of chemical transformations usually involving breaking and making of covalent bonds → prediction of new reactions! The power of unencumbered synthesis Biology - Chemistry Frontier Biology Intellectualization at the level of function Mechanisms: Identification of all components required to attain and maintain biological function → reconstitution! The power of bioreplicative synthesis Chemistry Intellectualization at the level of structure Mechanisms: Pathways of chemical transformations usually involving breaking and making of covalent bonds → prediction of new reactions! The power of unencumbered synthesis Biology - Chemistry Frontier On the Value of Natural Products in Pharma Discovery The de novo discovery of a new molecular agent of value in medicine is a daunting task. The risks are virtually incalculable. The study of small molecule natural products (SMNPs) may allow for entry into the discovery progression at a more advanced stage than the screening of medicinal chemistry sample collections, let alone standard diversity libraries. If properly exploited, this advantage may well compensate for the difficulties of exploration, collection maintenance, isolation, proof of structure and more complicated follow up synthesis studies which are often cited against the natural products based discovery route. Highly Successful Small Molecule Natural Product Derived Blockbuster Drugs (Partial List) (a) Antibiotics: β-lactam, aminoglycosides, macrolides (b) Statins: Zocor, Pravastatin, Lipitor (c) Steroids: Population control, antiinflammatories, cardiotonics, antiestrogens, antiandrogens, dermatological applications, etc. (d) Alkaloids: Antidepressants, antitumor agents (Vinca, thecins) (e) Terpenoids: Taxol (f) Polyketies: Anthracyclines Natural Products as Lead Candidates: The Statin Class of Antilipidemic Drugs HO HO O O H O O O O H HO O F N O CO2H OH O NHPh Lovastatin (Mevacor®) Natural Product (Isolated from A. terreus) Simvastatin (Zocor®) Natural Product-Derived Atorvastatin (Lipitor®) Natural Product-Inspired Why Have Natural Products Been a Rich Source of Drug Discovery? (a) Wisdom of the ages: While the developmental justifications for the biosynthesis of secondary metabolites are often mysterious, in many instances there could well be some overall rationale which can be exploited. (b) The small molecule natural product has possibly been designed for its ability to bind to biomacromolecules. (c) Presumably the biosynthesis of the natural product was under enzymatic control. At its “launching,” the natural product had been contained and extruded from some protein pocket. (d) There is often a built in advantage in that the natural product has already been maintained in some living host, without unmanageable toxicity. Why Chemical Synthesis of Complex Molecules? 1. The sheer challenge! 2. A setting for the evaluation of the scope of current methodology. 3. A setting for the evaluation of “problem areas”. 4. Incitement to strategy level and new methodology centered creativity. 5. A medium for the discovery and development of new drug possibilities through diverted total synthesis. 6. To provide material for pre-clinical and clinical investigation. 7. Molecular editing and diverted total synthesis. Diverted Total Synthesis Allow for the exploration of chemical space not available directly from the natural product. Through the process of Diverted Total Synthesis, it is possible to manipulate chemical functionality that can not be altered through manipulation of the natural product itself. DTS Starting Materials Analogs add complexity Chemical Synthesis Total Synthesis Natural Product reduce complexity Biosynthesis Advanced Intermediate DTS Analogs DTS = DIVERTED TOTAL SYNTHESIS Summary of the Epothilone Program: Molecular Editing through Total Synthesis S N O O Me O OH O OH S N H O O OH S N H Remove Epoxide Decrease toxicity Me Install Unsaturation Improve Potency and Biological Stability Me O O OH O OH O OH Epothilone B (EpoB) dEpoB (KOS-862) Phase II 9,10-dehydro-dEpoB (KOS-1584) Phase I S O N Install Trifluoro Group Decrease Toxicity and Broaden Therapeutic Index F3C H N O O OH Modify Heterocyclic Sector Increase Efficacy and Stability F3C H O O OH O OH O OH Fludelone (KOS-1591) Preclinical Iso-Fludelone (KOS-1803) Preclinical Iso-Fludelone: Therapy of Extra Large Tumors Day 25 Day 39 Therapeutic effect of Iso-fludelone against extra-large MX-1 tumors (30 mg/kg, Q12Dx4, 6hr-infusion, N=4) Day 53 The Radicicol Program Radicicol and Cycloproparadicicol Mode of Action: Potent inhibition of Hsp90. The first total synthesis of radicicol was completed in our laboratory in 2001. As reported, radicicol was confirmed to be a potent inhibitor of the molecular chaperone, Hsp90. Radicicol was largely inactive in mouse xenograft studies. We suspected that the epoxide moiety was responsible for the failure to translate protein inhibition to in vivo efficacy. Cycloproparadicicol was synthesized in our laboratory and shown to be an effective inhibitor of Hsp90. Importantly, this fully synthetic analog is active in in vivo settings. O H O OH Cycloproparadicicol H O HO Cl O H O H O HO O Cl OH Radicicol Isolated from M. nordinii Total Synthesis of Cycloproparadicicol H O H Me OTBS + HO Me H 1) n-BuLi; CO2 2) PPh3, DIAD 47% (2 steps) OTBS O Me H Me Co2(CO)8 100% TMSO H O O Me (CO)3Co (CO)3Co H OTBS Me 1) Grubbs' 2nd gen. cat. 57% 2) I2, 69% Ring-Closing Metathesis H O HO O Me H OTBS OH OH Cycloproparadicicol HO O O Me Cl H O Me Me H O O Me H OTMS 140єC, then silica gel 75% Diels-Alder Cycloaddition H OTBS Me Me The Radicicol Program Radicicol and Cycloproparadicicol Total Synthesis of the Tumor Cell Migration Inhibitor Migrastatin O O NH O O OMe O OTMS OMe H2O O O O Migrastatin OH OMe OMe O O O HO O NH O OTBS OMe OMe OH Diverted Total Synthesis: Migrastatin Wound Healing Assay No Serum Serum Serum + 200 nM Core O O Serum + 200 nM Migrastatin O O O O NH O OH OMe Migrastatin Core OMe Migrastatin OH Diverted Total Synthesis: Migrastatin Inhibition of Metastasis in Mouse Breast Tumor (4T1) Model O NH O OH OMe Lactam OMe Ketone OH Tumor size on day 21 16 14 Diameter of tumor (mm) 12 Number of colonies 10 8 6 4 2 0 Ke to ne (1 0m g/ kg ) Ke to ne (2 0m g/ kg ) La ct am (1 0m g/ kg ) La ct am (2 0m g/ kg ) co nt ro l Clonogenecity Assay 900 800 700 600 500 400 300 200 100 0 -100 Ke to ne (1 0m g/ kg ) Ke to ne (2 0m g/ kg ) La ct am (1 0m g/ kg ) La ct am (2 0m g/ kg ) co nt ro l Anti-Metastatic Activity of Migrastatin Analogs (MDA-MB-231 cells) O 3 O O Me O Me O Me Serum OH OMe HN O Me OH OMe Me Migrastatin Pre-surgery: 2,3-dihydro-migrastatin ether (ME) Serum + 10µM ME ME (pre) control group No Serum Post-surgery: control group ME (post) ME (pre+post) Collaboration with Moore Lab at MSKCC Migrastatin - Isomigrastatin Family Isolated from S. Platensis Me O O Me OH O OMe Isomigrastatin 3 O NH O O HN O 3 O O Me O Me OH OMe Migrastatin Isolation Isomigrastatin: Woo, E. J.; Starks, C. M.; Carney, J. R.; Arslanian, R.; Cadapan, L.;Zavala, S.; Licari, P. J. Antibiot. 2002, 55, 1411. Migrastatin: (a) Nakae, K.; Yoshimoto, Y.; Sawa, T.; Homma, Y.; Hamada, M.;Takeuchi, T.; Imoto, M. J. Antibiot. 2000, 53, 1130. (b) Nakae, K.;Yoshimoto, Y.; Ueda, M.; Sawa, T.; Takahashi, Y.; Naganawa, H.;Takeuchi, T.; Imoto, M. J. Antibiot. 2000, 53, 1228. Instability of Isomigrastatin OMe Me R O O O HO Me OMe HO Me O OH HO O OMe H2O SN2’ Me R Dorrigocin A epi-Dorrigocin A H2O H2O Isomigrastatin additionelimination Me R O HO Me OH HO Dorrigocin B O R= O O NH OMe water-assisted 3,3rearrangement HO Me O R O Ju, J.; Shen, B. et. al. JACS 2005, 1622 Migrastatin O Me Unsuccessful Early Strategies to Isomigrastatin O O Me R OP OMe O O Me R OP OMe (R) (R) trans-trans-dienolide R O O Me R OP OMe O Me O OP OMe 2,3-cis-6,7-trans-dienolide Total Synthesis of Isomigrastatin – Fragment Union OMe O + OTMS OMe Me OMe Me O O O NH O Ph O NB O Me Me H Ph 1) Ph3P 2) H 2, Pd/C PPh3 Ph3P B O O O NH =R O O OMe Me OMe Me A O OH MOMO O O BH3 O A + B Me O OMOM OMe R R LiMeCuCN Me HO Me R OTBS OMOM OMe OTBS OMOM OMe Krauss, I. J., Mandal, M. , Danishefsky, S. J. Angew. Chem. Int. Ed. Engl. 2007, ASAP Delayed Implementation of Dienolide Core – Completion of Synthesis (±) O OH PhSe 3 Me HO Me TBSO OMOM OMe Me O Me R OTBS OMOM OMe O PhSe O Me Me R OH O OMe O =R NH O xs. O PhSe EDCI/DMAP kinetic r esolution >8:1 d.r. at C2 Me O PhSe Grubbs II O Me OH O OMe R mCPBA either C2 epimer O O Me Me R OH O OMe cat. PMe3 Me O O Me OH OMe R O 21 % trans 36 % cis (separable) (+)-Isomigrastatin Krauss, I. J., Mandal, M. , Danishefsky, S. J. Angew. Chem. Int. Ed. Engl. 2007, ASAP Total Synthesis of Grandisine A (background) H • displays affinity for the human δ-opioid receptor (2.7 µM). • µ and κ agonist have been associated with adverse side effects when administered. • in animal models, δ-opioid agonists have been well tolerated. O O H O H H H 8 9 N γ-pyridone Grandisine A Grandisine A 8-epi 9-epi rel. energy 0 kcal/mol ~-6 kcal/mol ~-4 kcal/mol J. Org. Chem., 2005, 70, 1889 Total Synthesis of the Grandisine A Nuc.. E+ OH NCbz H NCbz O H NCbz H3C OTIPS H NCbz + O O O OTIPS BF3OEt2 -78 oC endo H ax. OTIPS NCbz + O NCbz TBAF AcOH/THF H H ax. NCbz eq. H H H O H O H OTES H O H ~5-10% from opposite face H O H H H O H NCbz O H O H O H H N O H OSi(Et)3 HO LHMDS (+)-Grandisine A unpublished results Carbohydrate-Based Antitumor Vaccines: Rationale • • Cell surface (tumor) antigens are carbohydrate structures commonly expressed in glycoproteins, mucins, and glycosphingolipids. Antibodies that recognize these glycoconjugates can, on occasion, be found in human sera. It has been postulated that these antibodies are part of an immune response to the tumor state. Antibody formation is provoked by suitable glycoconjugates and not by the oligosaccharides alone. Having been elicited in this way, most antibodies are primarily sensitive to the structure of the carbohydrate domain. Vaccination with carbohydrate vaccines may provide immunological protection against micrometastases and circulating tumor cells. The possibility of an immune response to cancer could have an enormous impact on the in vivo diagnosis and therapy of cancer. • • • Oligosaccharide Synthesis: How We Got Started OMe R1 R3SiO R2 O R3 OMe Lewis Acid R1 R3SiO R2 O R3 R1 HO R2 O R3 HO O HO HO OH OH HO O HO HO Classic Approach Glycal Assembly Approach Oligosaccharide Synthesis: The Glycal Assembly Method O O RO RO O O OH RO RO O O RO RO O O OH RO RO O O OH RO RO RO Oligosaccharide RO glycal RO RO O RO RO O RO P RO O P O RO O P HO O RO O O O O O OH RO O RO X O OH O O O O O OH XR O O RO RO E glycosyl donor RO O OR' E E+ E+ RO in situ generated glycosyl donor O + O I O I+ O PhSO2NH2 I O NHSO2Ph SEt I+ O NHSO2Ph O HO HO I O O I O O O I O O O O O NSO2Ph O O NHSO2Ph Carbohydrate Cancer Vaccine Program Angew. Chem. Int. Ed. 1996, 35, 1380 Breast Cancer-Associated Antigen: MBr1 Antigen O O O OTIPS AcO O O O OTIPS O SEt OBn OBn HO O BnO O OBn O O BnO BnO OBn OBn O O O NHSO2Ph O OBn OBn OBn HO HO OH O O O HO O OH O AcNH HO O OH O HO O HO OH O OH O HO OH O OH O O C25H51 HN C13H27 OH OH OH HO Globo-H Clinical Trials with Carbohydrate-Based Antitumor Vaccines HO HO OH O O O OH HO OH HO O OH O HO OH O HO O HO OH O OH O OH HO HO O NHAc OH O OH HO O OH O AcHN O AcHN OH HO HO OH O OH H N HO O O OH O AcHN O N H Linker O KLH O OH HO O Linker KLH Globo-H SCLC (Phase I) Prostate (Phase I) Breast (Phase I) OH O HO HO OH O O O OH HO HO HO Me OH OH O O O OH OH OH O O AcHN O O OH HO HO NHAc HO O O AcHN O HO O OH HO HO O OH TF(c) Prostate (Phase I) OH OH O HO OH O OH OH O O O NHAc O Linker KLH OH LewisY Ovarian (Phase I) HO HO OH O AcHN O HO HO H N O AcHN O O OH O AcHN O N H Linker O KLH O OH O OH HO2C O OH HO OH O OH O Linker KLH AcHN Fucosyl-GM1 SCLC (Phase I) HO HO O OH Tn(c) Prostate (Phase I) Flexible Accesses to Glycosyl-amino Acids pentenyl glycosides Protected carbohydrate allyl and pentenyl glycosides X Protected carbohydrate glycosyl donnors Protected carbohydrate O X = CH2 X=O CO2R O n X X = Glycosyl Donor CO2Bn NHFmoc HO CO2Bn NHFmoc O MeO P MeO NHBoc Horner -Emmons Protected carbohydrate Cross Metathesis CO2R NHBoc Protected carbohydrate Glycosylation (L.A.) CO2Bn NHFmoc Protected carbohydrate O O n O CO2Bn NHFmoc H2 (S, S)-Et-DuPHOS-Rh H2/Pt-C H2/Pt-C Protected carbohydrate O n CO2Bn NHFmoc n = 1 or 3 Unimolecular Multivalent Vaccine for Prostate Cancer: A Construct Containing Five Different Antigens OH OH O HO O Me HO O OH OH OH OH OH O O O O NAc HO O H OH HO Globo-H HO HO OH AcN H OH OH O OH HO O HO HO O OH O N H O N H O CO2H STn O HO O O AcHNO HO HO OH O AcHN O Tn HAcN HO HO O OH AcHN O O O HO2C OH HO O HO HO HO OH NHAc GM2 OH O O OH O OH HO HO O H N H N O H N O Linker KLH OH O O OH AcHN O TF HO O OH A fully synthetic, unimolecular pentavalent vaccine incorporating five of the known antigens associated with breast cancer has been synthesized and evaluated in preclinical settings. This compound is scheduled to enter Phase I clinical trials at MSKCC in fall / winter 2007. Erythropoietin (EPO): A Multiply Glycosylated Protein 126 38 161 Cysteine pair Cysteine pair 7 24 Sialic acid Galactos e annose M 33 83 29 Fucose N-Acetylglucosamine 166-Residue glycoprotein N-Glycosylated at Asn24, Asn38, and Asn83 O-Glycosylated at Ser126 Two cysteine pairs: Cys7-161 and Cys29-33 Heterogeneous glycoprotein Treatment of anemia Various glycoforms have different biological properties Hematopoietic growth factor Erythropoietin (EPO): A Multiply Glycosylated Protein EPO is a heterogeneous glycoprotein that may exist as a number of glycoforms. The amino acid sequence is highly conserved among isoforms. Current manufacturing processes yield inseparable mixtures of glycoforms, which can lead to complications at the regulatory level. Could we use total synthesis to gain access to single glycoforms of erythropoietin? We elected to synthesize the EPO isoform containing the consensus amino acid sequence and displaying highly branched and sialidated carbohydrate sectors. Incorporation of these types of carbohydrates is known to convey biostability. We will evaluate whether our fully synthetic erythropoietin is able to fold properly and whether it possesses EPO-like biological activity. The successful completion of this undertaking would require the development of some new synthetic methodologies to allow for the preparation and merging of very large and complex glycopeptide fragments. Toward Erythropoietin (EPO): The Development of Novel Methods for Glycopeptide Ligation Glycan O Peptide1 X H2N R Peptide2 O Glycan The Challenge Glycopeptide Ligation Glycan Peptide1 O R N H O Glycan Peptide2 I. A Cysteine-Based Glycopeptide Ligation Glycan O Peptide1 SEt S O StBu S H2N Peptide2 O Glycan Glycan O Peptide1 O Glycan O Peptide1 S HS H2N O Peptide2 Glycan HO H2N O HS Peptide1 O S Peptide2 Glycan Peptide1 Glycan Glycan O HS N H O Glycan Peptide2 Reduce S-S Toward Erythropoietin (EPO): The Development of Novel Methods for Glycopeptide Ligation II. A Cysteine-Free Ligation Model Glycan O Peptide1 P O S O H2N Glycan R Peptide2 O Glycan Glycan O Peptide1 P O S HN R Peptide2 O Removing P Glycan Glycan O Peptide1 R Glycan Glycan S Peptide2 O O Peptide1 R O O HO S HN SH HN O Peptide2 S Glycan N Glycan Glycan Glycan O Peptide1 R O Peptide2 O Peptide1 R N SH N H Peptide2 O HO Dr. Gong Chen Toward Erythropoietin (EPO): The Development of Novel Methods for Glycopeptide Ligation III. A Cysteine-Free Glycopeptide Ligation SEt Glycan O Glycan SP O H2N MeO OMe OMe MeO OMe OMe Glycan O Peptide1 SH N O R Peptide2 Glycan R Peptide2 O Peptide1 S O Glycan R Philip Dawson R'S S HN O Peptide2 Glycan O Peptide1 S HN O R Glycan Peptide2 MeO OMe OMe Glycan O Peptide1 N H R Peptide2 O MeO OMe OMe Glycan Toward Erythropoietin (EPO): The Development of Novel Methods for Glycopeptide Ligation IV. A Cysteine-Free Glycopeptide Condensation SEt S O H2N O Glycan R Peptide2 Glycan O Peptide1 Saburo Aimoto AgCl HOOBt, DIEA DMSO, RT Gly / Pro Glycan O Peptide1 N H R Glycan Peptide2 O Lys and Cys Protected SEt S O H2N O Glycan R Peptide2 O S O O Glycan O Peptide1 Gly / Pro TCEP HOOBt, DIEA DMSO, RT Glycan O Peptide1 N H R Glycan O Peptide2 O S O O Lys and Cys Protected Toward Erythropoietin (EPO): The Development of Novel Methods for Glycopeptide Ligation V. Free Radical Desulfurization: Cys to Ala Glycan O Peptide1 N H Peptide2 O SH Glycan TCEP, tBuSH, VA-50 H2O, RT Glycan O Peptide1 N H Glycan Peptide2 O S-H Peptide In . . S Peptide R P R R S Peptide . R P R R . CH2 Peptide H-S CH3 Peptide R S P R R P(CH2CH2COOH)3 TCEP O FmocN S Me S Acm S S O O HN N N H2N NH HCl NH2 tolerated VA-50 Progress Toward Erythropoietin (EPO): Synthesis of Glycan BnO OBn O OBn F BnO OBn O OBn O O O BnO PhSO2NH OTBS BnO O AcO BnO PhSO2NH SEt TIPSO HO BnO O BnO O AcO BnO PhSO2NH HO O BnO O BnO O HO BnO PhSO2NH Ph O O PMBO OBn O S BnO BnO BnO Ph BnO OBn O OBn BnO O O O BnO PhSO2NH OBn O O O BnO PhSO2NH OTBS OBz O O 2x SEt OH BnO BnO BnO O O BnO BnO BnO BnO O BnO OBn O OBn BnO O O O BnO PhSO2NH OBn O O O BnO PhSO2NH OTBS HO BnO HO O OH Progress Toward Erythropoietin (EPO): Synthesis of Glycan O O O OBn O + HO BnO OBn O O O O OBn O BnO O BnO OBn O HO HO OBn O BnO OBn O SEt O BnO OBn O SEt O O TMSESO2NH2 EtSH AcO OAc O AcO PivO O OBn + HO BnO O SEt CCl3 NH NPht AcO OAc O O AcO BnO PivO NPht NPht AcO OBn O BnO O SEt OAc OAc O OP(OBn)2 CO2Me AcO HO HO OBn O BnO AcHN AcO CO2Me AcO OBn O O O AcHN O BnO AcO OBn OAc OAc OBn O SEt NPht NPht Progress Toward Erythropoietin (EPO): Synthesis of Glycan OH CO2Me AcO OAc OBn AcO OBn OAc O O O O AcHN O BnO AcO OBn PhtN BnO BnO BnO O O SEt BnO OBn O OBn BnO O O O BnO PhSO2NH OBn O O O BnO PhSO2NH OTBS + BnO BnO BnO BnO O 2x O OH AcO CO2Me OBn AcO OBn O O O O AcHN O BnO AcO OBn PhtN BnO BnO BnO OAc OAc O BnO O O BnO O OBn O OBn AcO CO2Me AcO OBn BnO O O O O AcHN O BnO AcO OBn PhtN OAc OAc BnO BnO BnO OBn BnO O O O BnO PhSO2NH O 1. O O 2. O BnO PhSO2NH OTBS Global Deprotection Kochetkov Amination O Progress Toward Erythropoietin (EPO): Synthesis of Glycan HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN HO HO HO OH OH O OH OH O OH O HO O HO O AcHN O OH O AcHN NH2 O O HO O O HO O HO CO2H OH OH HO O O O O AcHN O HO HO OH AcHN OH OH HO HO HO O Progress Toward Erythropoietin (EPO): Synthesis of Glycan HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN OH OH HO HO HO O O O OH O HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN OH OH HO OH OH HO HO O HO O HO O HO O AcHN HO O HO O AcHN NH2 O CO2H OH OH HO O O O O AcHN O HO HO OH AcHN O Progress Toward Erythropoietin (EPO): Synthesis of Glycan OAc OAc AcHN AcO OAc OAc AcHN AcO AcO AcO OAc OAc AcHN AcO O O O O OAc HO N3 AcO OAc OAc AcHN AcO AcO OAc OAc AcHN AcO FmocHN O OBn O O O O OAc OAc O O NHAc O OAc O O COOMe O FmocHN O O OBn OAc O SEt O O O O OAc OAc AcHN AcO OAc OAc AcHN O AcO OAc O COOMe O O OP(OBn)2 COOMe O O N3 O O OTIPS O O O N3 OH O COOBn NHFmoc OTIPS O O CCl3 NH AcO OP(OBn)2 COOMe HO OH OTIPS O O O OTIPS O AcO AcO Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN HO HO HO OH OH O O O HO O OH O OH OH O HO O HO O AcHN O O OH OH O AcHN NH2 O O HO HO HO CO2H HO OH OH HO HO O O O O AcHN O HO HO OH AcHN OH OH + O Ala Glu Asp Ile Dmab O Thr Thr Gly O SSEt Aspartylation HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN HO HO HO OH OH O O O HO O OH O OH OH O HO O HO O AcHN O Ala Glu Asp Ile O Dmab OH O AcHN O Thr Thr Gly O SSEt H2N Cys Ala Glu His Cys Ser Leu Asn Glu H N O HS O O HO CO2H HO OH OH OH OH HO O O O O AcHN O HO HO OH AcHN HO HO HO NCL Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN HO HO HO OH OH O O O HO O OH O OH OH O HO O HO O AcHN O Ala Glu Asp Ile O Dmab OH O AcHN Thr Thr Gly Cys Ala Glu His Cys Ser Leu Asn Glu H N O O HO O HO CO2H OH OH HO O O O O AcHN O HO HO OH AcHN OH OH HO HO HO EPO 22-37 Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN HO HO HO OH OH O O O HO HO HO CO2H HO OH OH HO HO O O O O AcHN O HO HO OH AcHN OH OH O OH O HO O HO OH OH O O AcHN OH O AcHN NH2 O O HO + OH O O Thr Thr Gly S O O O O Aspartylation Removing Fmoc FmocHN Lys Glu Ala Glu Asp Ile Allyl ivDde Allyl HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN HO HO HO OH OH O O O HO O OH O HO O HO OH OH O O AcHN OH O AcHN H N O O HO O O O S O HO CO2H HO OH HO OH OH HO OH HO O O O O AcHN O HO HO OH AcHN O O H2N Lys Glu Ala Glu Asp Ile Allyl ivDde Allyl Thr Thr Gly TCEP Fragment Condensation + O O SSEt Ala Pro Pro Arg Leu Ile Cys Asp Ser Arg Val Leu Glu Arg Tyr Leu Glu Ala Acm Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN HO HO HO OH OH O O O HO O OH O OH OH O HO O HO O AcHN O O OH O AcHN H N O O O HO HO CO2H OH OH HO O O O O AcHN O HO HO OH AcHN OH OH HO HO HO ? Thr Thr Gly Allyl ivDde Allyl O S O O Ala Pro Pro Arg Leu Ile Cys Asp Ser Arg Val Leu Glu Arg Tyr Leu Glu Ala Lys Glu Ala Glu Asp Ile Acm EPO 1-28 Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks AcO OAc OAc O AcO OAc OAc O COOMe O AcHN AcO OAc OAc O AcO O O O OAc O O AcNH O Ala Ala Asp Pro Pro Ser Ile Ser Ala SSEt O O SH HN Ala Pro Leu Arg Thr Ile ivDde Ala Glu Lys Gln Ala NHFmoc ivDde AcHN Auxiliary Ligation Thr Ala Asp Thr Phe Arg Lys Leu Phe Arg Val Tyr Ser Asn + MeO OMe OMe Arg Asp Gly Thr Arg Cys Ala Glu Gly Thr Tyr Leu Lys leu Lys Gly Arg Leu Phe Acm ivDde ivDde Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks AcO OAc OAc O AcO OAc OAc O COOMe O AcHN AcO OAc OAc O AcO O O O OAc O O AcNH O Ala Ala Asp Pro Pro Ser Ile Ser Ala Ala Pro Leu Arg Thr Ile Thr Ala Asp Thr Phe Arg Lys Leu Phe Arg Val Tyr Ser Asn Arg Asp Gly Thr Arg Cys Ala Glu Gly Thr Tyr Leu Lys leu Lys Gly Arg Leu Phe Acm ivDde ivDde Ala Glu Lys Gln Ala NHFmoc ivDde AcHN ivDde EPO 114-166 Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks AcO OAc OAc O AcO OAc OAc O COOMe O AcHN AcO OAc OAc O AcO O O O OAc O O AcNH O Ala Ala Asp Pro Pro Ser Ile Ser Ala Ala Pro O SSEt O H2N-- Leu Arg Thr Ile ivDde Ala Glu Lys Gln Ala NHFmoc ivDde AcHN TCEP Fragment Condensation Thr Ala Asp Thr Phe Arg Lys Leu Phe Arg Val Tyr Ser Asn Arg Asp Gly Thr Arg Cys Ala Glu Gly Thr Tyr Leu Lys leu Lys Gly Arg Leu Phe Acm ivDde ivDde Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks AcO OAc OAc O AcO OAc COOMe O AcHN AcO OAc OAc O AcO O O O OAc OAc O O O AcNH O Ala Ala Asp Pro Pro Ser Ile Ser Ala Ala Pro Leu Arg Thr Ile Thr Ala Asp Thr Phe Arg Lys Leu Phe Arg Val Tyr Ser Asn Arg Asp Gly Thr Arg Cys Ala Glu Gly Thr Tyr Leu Lys leu Lys Gly Arg Leu Phe Acm ivDde ivDde Ala Glu Lys Gln Ala NHFmoc ivDde AcHN ivDde EPO 114-166 Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN HO HO HO OH OH O O O HO HO HO CO2H HO OH OH HO HO O O O O AcHN O HO HO OH AcHN OH OH O OH O OH OH O HO O HO O AcHN O SSEt O O O OH O AcHN O O HO OH O + NH2 Ser Ser Asp Val Leu Leu Ala Gln NHFmoc Gln Pro HO Aspartylation CO2H OH OH OH O O O O AcHN O HO HO OH AcHN HO HO HO OH OH O O O HO O OH O OH OH O HO O HO O AcHN O Ser Ser Asp Val Leu Leu Ala Gln NHFmoc O Gln SSEt O O Pro OH O AcHN H N O O HO O HO CO2H OH OH HO O O O O AcHN O HO HO OH AcHN OH OH HO HO HO Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN HO HO HO OH OH O OH OH O OH O HO O HO O AcHN O Ser Ser Asp Val Leu Leu Ala Gln NHFmoc O SSEt O O Gln Pro OH O AcHN H N O O HO O O HO O TCEP Fragment Condensation O HO CO2H OH OH HO O O O O AcHN O HO HO OH AcHN OH OH HO HO HO + H2N Trp Glu Pro Leu Gln Leu His Val Asp Lys Ala Val Ser Gly Leu Arg Ser O O S O Thr Gly Leu Ala Arg Leu Leu Thr Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks HO CO2H OH OH OH O O O O AcHN O HO HO OH AcHN HO HO HO OH OH O HO O O HO O OH O OH O OH O AcHN Ser Ser Asp Val Leu Leu Ala Gln NHFmoc H N O O HO HO O HO O AcHN O HO CO2H OH OH HO O O O O AcHN O HO HO OH AcHN OH OH HO HO HO O O Gln Pro Trp Glu Pro Leu Gln Leu His Val Asp Lys Ala Val Ser Gly Leu Arg Ser O O S O Thr Gly Leu Ala Arg Leu Leu Thr EPO 78-113 Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks O O S O EPO 78-113 Gly HO CO2H OH OH OH E O O O O AcHN O HO HO OH AcHN HO HO HO OH OH NH2- Ala EPO 114-166 O OH OH O OH O HO O HO O AcHN O OH O AcHN H N O O HO O O HO O AgCl Fragment Condensation O Saburo Aimoto HO CO2H OH OH HO O O O O AcHN O HO HO OH AcHN OH OH HO HO HO O Ser Ser Asp Val Leu Leu Ala Gln NHFmoc Gln AcO OAc OAc O AcO OAc Pro Trp Glu Pro Leu Gln Leu His Val Asp Lys Ala Val Ser Gly Leu Arg COOMe O Thr Ala Ala Asp Pro Pro Ser Ile O Ser Ala Glu Lys Gln Ala Gly Leu Ala Arg Leu Leu Thr ivDde Ser AcHN AcO OAc OAc O AcO O O O OAc OAc O AcHN O AcNH O ivDde Ala Ala Pro Leu Arg Thr Ile Thr Ala Asp Thr Phe Arg Lys Leu Phe Arg Val Tyr Ser Asn Arg Asp Gly Thr Arg Cys Ala Glu Gly Thr Tyr Leu Lys leu Lys Gly Arg Leu Phe Acm ivDde ivDde EPO 78-166 Progress Toward Erythropoietin (EPO): Assembly of the Glycopeptide Building Blocks Lys leu Lys Leu Tyr Thr Gly Glu Ala Cys Arg Thr Gly Asp Arg Gly Arg Leu Ala Pro Pro Arg Leu Ile Phe Asn Ser Tyr Asp Val Arg Phe Leu Lys Arg Phe Thr Asp Ala Thr Ile Thr Arg Leu Pro Ala Ala Ser Ala Ala Asp Pro Pro Ser Ile Ala Glu Lys Ala Val Ser Gly Leu Arg Ser Leu Thr Thr Leu Leu Arg Ala Leu Gly Ala Gln Lys Asp Val His Leu Gln Leu Pro Glu Trp Pro Gln Ser Ser Asn Val Leu Leu Ala Gln Gly Arg Leu Lys Val Ser Trp Lys Arg Met Glu Val Gly Gln Gln Ala Val Glu Trp Gln Gly leu Ala leu leu Ser Glu Ala Val Lys Thr Asp Pro Val Thr Ile Asn Glu Asn Leu Ser Cys His Glu Ala Cys Gly Thr Thr Ile EPO 1-28 Cys Asp Ser Arg Val Leu Glu Arg Tyr Leu Glu Ala Lys Glu Ala Glu Asn EPO 29-77 EPO 78-166 Summary We close this review with some thoughts concerning life on the chemistry– biology frontier. In this account, we have shown by historical progression how our laboratory, starting with fascinating problems in the field of “small molecule” natural products, has become involved in issues of tumor expression and tumor immunology. One of the singular contributions which we and like-oriented research groups bring to such coalitions is a sensitivity for precisely defined structures. When collaborating with biologists in identifying bioactive compounds and charting their functions, the chemist insists that the compounds in question be demonstrably pure and that the structural assignments, down to each stereogenic center, be corroborated. But chemistry’s contribution to the enterprise is certainly more than restraints arising from insistence on thoroughness and intellectual exactitude. Methodological building upon the principles of our science leads to the magic of synthesis – with its unique capability to prepare molecules of virtually any shape and juxtaposition of functional groups. Creative synthesis is the indispensable talent that the chemist will bring to the many exciting struggles and opportunities in the future. Angew. Chem. Int. Ed. 1996, 35, 1380 Thank you!

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