CURRICULUM VITAE

NAME:          Ravi Allada

BIRTH DATE: August 20, 1967

EDUCATION: University of Michigan           Integrated Premedical Medical Program (Inteflex)
           Ann Arbor, Michigan              BS (With Highest Distinction) in Biomedical Sciences,
                                            MD (With Distinction and Distinction in Research),

  Assistant Professor
  Department of Neurobiology & Physiology
  Northwestern University
  2205 Tech Drive, Evanston, IL 60208-3520
  Phone: 847-491-2809 - E-mail:

 1990-1992    Howard Hughes Medical Institute-National Institutes of Health Research Scholar,
                 National Institute of Mental Health, Bethesda, MD (Advisor: Howard Nash)
 1994         Howard Hughes Medical Institute-National Institutes of Health Continued Support
                 Fellow, National Cancer Institute, Bethesda, MD (Advisor: Carl Wu)
 1994-2000    Clinical Pathology Resident, Department of Pathology, Brigham and Women’s
                 Hospital, Boston, MA (Chair: Ramzi Cotran)
 1995-2000    Postdoctoral Fellow, Department of Biology, Brandeis University, Waltham, MA
                 (Advisor: Michael Rosbash)
 2000-present Assistant Professor, Department of Neurobiology and Physiology, Northwestern
                 University, Evanston, IL

 2000-Present Assistant Professor, Department of Pathology, Feinberg School of Medicine,
                Northwestern University, Chicago, IL
 2000-Present Member, Robert H. Lurie Cancer Center, Northwestern University
 2000-Present Member, Center for Sleep and Circadian Biology, Northwestern University, Evanston,
 2002-Present Associate Director, Center for Sleep and Circadian Biology, Northwestern University,
                Evanston, IL
 2000-Present Member, Northwestern University Institute for Neuroscience, Northwestern University
 2000-Present Member, Interdepartmental Program in Biological Sciences, Northwestern University,
                Evanston, IL
 2001-Present Member, Center for Genetic Medicine, Northwestern University
 2004-Present Assistant Professor, Department of Biochemistry, Molecular Biology, and Cell Biology,
                Northwestern University, Evanston, IL

  Class Honors, University of Michigan, 1985-1989
                                                                                    Curriculum Vitae
                                                                                         Ravi Allada
  American Urological Association Student Fellowship, 1987
  American Heart Association Summer Student Research Fellowship, 1988, 1989
  Medical School Fund Scholarship, 1988
  Phi Beta Kappa, 1989
  Lange Medical Publications Award, 1989
  Dean’s Award for Academic Excellence, 1989, 1990
  Alpha Omega Alpha Honor Medical Society, 1993
  Howard Hughes Medical Institute-National Institutes of Health Research Scholar, 1990-1992
  Howard Hughes Medical Institute Fellowship for Continued Support, 1992-1994
  Hewlett-Packard Award, 1994
  National Research Service Award, 1995-1996
  Chief Resident, Clinical Pathology, Brigham and Women’s Hospital, 1996
  Howard Hughes Medical Institute Postdoctoral Research Fellowship for Physicians, 1996-1999
  Paul E. Strandjord Young Investigator Award, Academy of Clinical Laboratory Physicians and
   Scientists, 1998
 Burroughs Wellcome Career Award in the Biomedical Sciences, 1999-2003
 Whitehall Foundation Grant, 2000-2003
 National Alliance for Research on Schizophrenia and Depression Young Investigator Award, 2003-
 Burroughs Wellcome Fund Career Award Outsanding Progress Report, 2003
 March of Dimes Grant, 2004-present
 Presentation selected as “Hot Topic” at Chronobiology Gordon Research Conference (Valerie

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                                                                                       Curriculum Vitae
                                                                                            Ravi Allada



1.   Lin J-M, Schroeder A, and Allada R. (2005) In vivo circadian function of CK2 phosphorylation
     sites in Drosophila PERIOD. J Neurosci (in press).

2.   Lear BC, Lin J-M, Keath JR, McGill JM, Raman IM, and Allada R. (2005) The ion channel
     narrow abdomen is critical to the neural output of the Drosophila circadian pacemaker. Neuron
     (in press, article to be featured on the cover).

3.   Lear BC, Merrill CE, Lin J-M, Schroeder A, Zhang L, and Allada R. (2005) A novel G-protein
     coupled receptor, groom-of-PDF, is required for PDF neuron action in circadian behavior.
     Neuron (in press).

4.   Allada R, Kadener S, Nandakumar N, and Rosbash M. (2003) A recessive mutant of Drosophila
     Clock reveals a role in circadian rhythm amplitude. EMBO J 22: 3367-75.

5.   Zhao J, Kilman VL, Keegan KP, Ping Y, Emery P, Rosbash M, and Allada R. (2003)
     Drosophila Clock can generate ectopic circadian clocks. Cell 113, 755-766.

     Commentary on this paper: Hastings MH. (2003) Circadian clocks: self-assembling oscillators?
     Current Biology 13, R681-R682.

     Campbell N. (2003) Clockwork conductor. Nature Reviews Genetics 4, 578.

     Pilcher HN. (2003) Flies get rhythm. Nature 423, 935.

6.   Nash HA, Scott RL, Lear BC, and Allada R. (2002) An Unusual Cation Channel Mediates
     Photic Control of Locomotion in Drosophila. Current Biology 12, 2152-2158.

7.   Lin J-M, Kilman VL, Keegan K, Paddock B, Emery-Le M, Rosbash M, and Allada R. (2002) A
     Role for Casein Kinas            Drosophila Circadian Clock. Nature 420, 816-820. Advance
     online publication, 24 November 2002. (doi:10.1038/nature01235)

     Commentary on this paper: Blau J. (2003) A new role for an old kinase: CK2 and the circadian
     clock. Nature Neurosci 6, 208-210.

8.   Poteat HT, Chen P, Loughlin KR, Winkelman JW, Allada R, Ma'luf N, Tanasijevic MJ, Bates
     DW. (2000) Appropriateness of prostate-specific antigen testing. Am J Clin Pathol 113, 421-

9.   So WV, Sarov-Blat L, Kotarski CK, McDonald MJ, Allada R, Rosbash M. (2000) takeout, a
     novel Drosophila gene under circadian clock transcriptional regulation. Mol Cell Biol 20, 6935-

                                                                                               Page 3
                                                                                         Curriculum Vitae
                                                                                              Ravi Allada

10.   Allada R, White NE, So WV, Hall JC, Rosbash M. (1998) A mutant Drosophila homolog of
      mammalian Clock disrupts circadian rhythms and transcription of period and timeless. Cell 93,

      Commentary on this paper: Schibler U. (1998) Circadian rhythms: New cogwheels in the
      clockworks. Nature 393, 620 – 621.

11.   Wisniewski J, Orosz A, Allada R, Wu C. (1996) The C-terminal region of Drosophila heat
      shock factor (HSF) contains a constitutively functional transactivation domain. Nucleic Acids
      Res 24, 367-374.

12.   Allada R and Nash HA. (1993) Drosophila Melanogaster as a Model for Study of General
      Anesthesia: The Quantitative Response to Clinical Anesthetics and Alkanes. Anesth Analg 77,

13.   Terpenning MS, Allada R, and Kauffman CA. (1989) Intermittent Urethral Catheterization in
      the Elderly. J Am Geriatr Soc 37, 411-6.


1.    Allada R and Meissner R-A. (2005) Casein kinase 2, circadian clocks, and the flight from
      mutagenic light. Mol Cell Biochem 274, 141-149.

2.    Allada R. (2004) It’s About (Molecular) Time. Nat Cell Biol 6, 1161. Book review

3.    Allada R. (2003) Circadian Clocks: A Tale of Two Feedback Loops. Cell 112, 284-6.
             Invited Preview

4.    Turek FW and Allada R. (2002) Liver Has Rhythm. Hepatology 35, 743-5.

5.    Allada R, Emery P, Takahashi J, Rosbash M (2001) Stopping Time: The Genetics of Fly and
      Mouse Circadian Clocks. Annu Rev Neurosci 24, 1091-1119.

6.    Allada R and Rosbash M. (2000) Fly Clocks. In Genetic Models in Cardiorespiratory Biology,
      Haddad GG and Xu T, eds. (New York, NY: Marcel Dekker) pp. 365-390.

7.    Ishida N, Kaneko M, and Allada R. (1999) Biological Clocks. Proc Natl Acad Sci USA 96, 19-

8.    Rosbash M, Allada R, Dembinska M, Guo WQ, Le M, Marrus S, Qian Z, Rutila J, Yaglom J,
      Zeng H. (1996) A Drosophila Circadian Clock. Cold Spring Harbor Symp. Quant. Biol. LXI,
                                                                                                 Page 4
Curriculum Vitae
     Ravi Allada

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                                                                                            Curriculum Vitae
                                                                                                 Ravi Allada

                                  INVITED PRESENTATIONS

1.    First Annual Japanese-American Frontiers of Science Symposium, 1998, Irvine, California
2.    San Diego State University, Department of Biology, 1998
3.    New York University School of Medicine, Department of Biochemistry, 1998
4.    University of Chicago, Department of Pathology, 1998
5.    National Institute of Child Health and Development, Laboratory of Molecular Biology, 1999
6.    Northwestern University Medical School, Department of Pathology, 1999
7.    Signalling 2000, Keystone, Colorado, 2000
8.    Chicago Graduate Student Association Seminar Seies, 2001
9.    Molecular Cell Biology Gordon Research Conference, Tilton, New Hampshire, 2003
10.   University of Massachusetts Medical School, Department of Neurobiology, Worcester, MA, 2003
11.   Northern Illinois University, Department of Biological Sciences, DeKalb, Illinois, 2003
12.   University of California at Los Angeles, Neurogenetics Affinity Group Los Angeles, CA, 2004
13.   University of Southern California, Department of Molecular and Computational Biology, Los
      Angeles, California, 2004
14.   University of Pennsylvania, Department of Neuroscience, Philadelphia, PA, 2004
15.   4th International Conference on Protein Kinase CK2, University of Western Ontario, London,
      Ontario, 2004
16.   New York University, Department of Biology, New York, NY, 2004
17.   University of Missouri, Division of Biological Sciences, Columbia, MO May, 2005
18.   National Insitutes of Mental Health, Drosophila Neurobiology Interest Group, May 2005

                                      RESEARCH SUPPORT


National Institutes of Health, 1R01NS052903-01, “Function of the Novel Ion Channel na in Daily
       Rhythms,” 9/1/2005-8/31-2009, $925,000 total direct, $1,373,624 (total award)

National Allliance for Research on Schizophrenia and Affective Disorders Young Investigator Award,
       “Behavioral Genetics of a Drosophila Homolog of a Candidate Bipolar Susceptibility Gene,”
       7/1/03-11/30/05, $60,000 (total award)

National Institutes of Health, 1R01MH067870-01, “The Function of Casein Kinase 2 in the Circadian
       Clock,” 4/1/03-3/31/08, $1,150,000 total direct, 1,708,271 (total award)

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                                                                                         Curriculum Vitae
                                                                                              Ravi Allada
Burroughs Wellcome Fund Career Award in the Biomedical Sciences, #992844, “Molecular genetic
      analysis of the circadian rhythm gene, Clock, in Drosophila”, 9/1/99-8/31/06, $452,000 (total

March of Dimes, #1-FY04-106, “Analysis of the Circadian Rhythm Gene Clock in the Induction and
      Organization of MolecularPacemakers,” 6/1/04-5/31/07, $204,208 total direct, $224,629 (total


Department of Defense (subcontract from NuNetix), “Identification and Characterization of Molecular
      Inhibitors of Cognitive Performance”, $14,467, 1/1/05-6/30/05

DARPA, “Mutant Mice with Altered Sleep-Wake Phenotype: Molecular Pathways and Targets for Drug
    Development,” 4/1/04-12/31/06, $4,100,000 (total award), Turek (PI), Allada (co-PI), Bass,
    Dugovic, Laposky (co-PIs).

Whitehall Foundation Grant, #2000-08-53-APL, “The Cloning and Analysis of New Circadian Clock
      Genes,” 9/1/00-8/31/04, $211,760 total direct, $225,000 (total award)

Howard Hughes Medical Institute Postdoctoral Research Fellowship for Physicians, 9/1/96-8/31/99

National Institutes of Health Research Service Award, 7/1/95-8/31/96

Howard Hughes Medical Institute National Institutes of Health Research Scholar, 7/1/90-6/31/92

                                     OTHER ACTIVITIES

•    NIH National Center on Sleep Disorder Research Workshop on “Neurobiology of Sleep and
     Working: Implications for Insomnia”, Panel member, (09/01)

•       Member, Special Emphasis Panel for Neurogenesis and Cell Fate (NCF) Study Section (2005)

•       Member, Biological Rhythms and Sleep (BRS) Study Section (2005-present)

•       Ad Hoc, MDCN-3

•       Ad Hoc, NSF

•    Cell
•    Neuron
•    Current Biology
•    Molecular and Celluar Biology
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                                                                           Curriculum Vitae
                                                                                Ravi Allada
•      Journal of Neurophysiology

•    American Association for the Advancement of Science
•    Society for Reseach on Biological Rhythms
•    Society for Neuroscience

Research Assistant Professors:
Dr. Valerie Kilman      (Ph.D. Brandeis University)                2005-present

Postdoctoral Fellows:
Dr. Jui-Ming Lin        (Ph.D. Illinois Institute of Technology)   2001-present
Dr. Valerie Kilman      (Ph.D. Brandeis University)                2001-2004
Dr. Bridget Lear        (Ph.D. University of Chicago)              2002-present
Dr. J.Russel Keath      (Ph.D. University of Massachusetts)        2003-present
Dr. C. Elaine Merrill   (Ph.D. Vanderbilt University)              2004-present

Graduate Students:
Jena Pitman                         Ph.D.          2002-present
Natalie Boone                       M.S.           2002-2003
Kevin Keegan                        Ph.D.          2003-2005
Rose-Anne Meissner                  Ph.D.          2004-present
Luoying Zhang                       Ph.D.          2004-present
Jeff Guzelian                       M.S.           2005-present

Graduate Rotation Students/Undergraduate Students:
A. Rotation Students
Jayshree Biswas (NUIN, 2001)
Lisa Sharpe (IBIS, 2001)
Lisa McCarty (IBIS, 2002)
Jena Pitman (NUIN, 2002)
Lamaar Cunningham (NUIN, 2002)
Kristin Konigsfeld (IBIS, 2002)
Kevin Keegan (IBIS, 2003)
Georgette Moyle (IBIS, 2003)
Rose-Anne Meissner (NUIN, 2004)
Luoying Zhang (NUIN, 2004)
Xiao-Xiao Wang (IBIS, 2005)
Sandhya Rayappa (IBIS, 2005)
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                                                                                        Curriculum Vitae
                                                                                             Ravi Allada
Deanna Arble (NUIN, 2005)

B. C98 and C99 Independent Study Students:
Salman Khan                    2000
Rudi Scharnweber               2000-2001
Elyssa Burg (Macey Scholar)    2000-2003
Alia Majeed                    2001-2003
Dave Hanrahan                  2002-2004
Matt Krzmienski                2003-2004
Nelson Moy                     2003-2004
Suraj Pradhan                  2005-present

                                            COURSES TAUGHT

    •   *Genes, the Brain, and Behavior (NUIN 485)            (2001, 2004)
    •   Molecular Biology (BIOL SCI 390)                      (2001-present)

    •   Medical Scientist Training Program Research Seminar   (2001-2003)
          (Neurobiology Block)

    •   *NUIN Fundementals of Neuroscience Core Course        (2001-present)
        (Signaling, Molecular Biology lectures)

    •   *Sleep (NUIN)                                (2005)

    •   Functional Genomics, guest lecturer          (2002)

    •   IBIS Core course, guest lecturer             (2002)

    *Indicates development of new course/lectures

                                           UNIVERSITY SERVICE

•        Chair, Center for Sleep and Circadian Biology Journal Club (2001-present)
•        Member, IBIS Admissions Committee (2001-03)
•        Member, NUIN Graduate Student Advisory Committee (2001-03)
•        Member, Dunbar Scholar Faculty Search Committee (2001-03)
•        Member, Neurobiology and Physiology Seminar Committee (2002-present)
•        Member, MSTP Admissions Committee (2002-2004)
•        Member, BMBCB Faculty Search Committee (2002-03)
•        Member, Evanston Northwestern Healthcare-Northwestern University Joint Faculty Search
         Committee (2001-2002)
•        Member, Shared Facilities Committee (2003-present)
•        Member, IBIS Advisory Committee (2003-present)
•        Member, Robotics Facility Advisory Board (2004-present)
•        Associate Director, Center for Sleep and Circadian Biology (2004-present)
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                                                                                          Curriculum Vitae
                                                                                               Ravi Allada
•       Freshman Advisor, Honors Program in Medical Education (2003-present)


     Mentor, Evanston Township High School Science program (2001)

     Mentor, Illinois Math and Science Academy Internship program, Hussein Bhihkapurwala and
               Jonathan Delich (2003)

    Head Coach, Rookie League Dodgers, Evanston Baseball and Softball Association (2005)

    Chief Scientific Officer, NuNetix: founded with Northwestern colleagues (Fred Turek, Joe Bass,
        Christine Dugovic, Aaron Laposky, and Sue Olson) NuNetix, a company focused on high
        throughput genetic and genomic approaches to sleep, circadian rhythms and obesity (2004-

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