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(CELL CYCLE)

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(CELL CYCLE)

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(CELL CYCLE)

Phases of cell cycle

Every individual of a given species has a characteristic number of chromosomes in its body cells. The normal

chromosomal constitutions are called karyotypes.

Usually when cells reach a certain size, they must either stop growing or divide, Some cells such as nerve, skeletal

muscle, and red blood cells do not normally divide once they as nerve, skeletal muscle, land red blood cells do not normally

divide once they are mature. The actities of growing and dividing cells, can be described in terms of the life cycle of the cell

or the cell cycle. Such cycles start each time when new cells are formed, and they end when those cells compete their own

division. The cycle starts again for each new daughter cell. The process that occurs during the division of eukaryotic cells is

conveniently diagramed as a cells cycle:

G1 S G M C

Interphase

Most of the cell cycle is spent in interphase. Chromosomes duplicate during this period, many cell parts are made,

and the cell does most of its growing. Typically, intyerphases lasts for at least 90% of the total time required for the cell

cycle. DNA synthesis, the main event in chromosome duplication, occurs in the middle of interphase and serves as the basis

for dividing interphase into three sub-phases.

G1 phase

The firs subphase, designated G1 is the period befor DNA synthesis begins. G stands for gap and G 1 refers to the

firs gap between cell division and DNA synthesis. This phase is a gap between cell activity. In G 1 phase, the cell increases

its supply of proteins, increases the number of many of its organelles, such as mitochondria and ribosomes, and grows in

size. Towards the end of G1 phase there is an increased activity of enzymes required for DNA synthesis.

S Phase: Following G1 phase is the sub-phase S, in which DNA synthesis (replication) actually occurs. At the beginning of

the S phase, each chromosome is single. At the end of this phase, after DNA replication, the chromosomes are double, each

consisting of two sister chromatids.

G2 Phase: The third sub-phase, called G2 phase, spans the time from the complition of DNA synthesis to the onset of cell

division. In this phase, increased protein synthesis occurs as the final step in the cells preparation for cell division. The

completion of G2 phase in marked by the beginning of mitosis. The sequence of the sub-stages of interphase is therefore:

G1 phase S Phase G2 Phase

M phae: In this the micro-tubular apparatus is assembled, binds to the chromosomes, and moves the siter chromosomes

apart. This phase is called mitosis.

C phase: In this phase the cell itself divides, creating two dauthter cells. This phase is called cytokinesis.

Length of each phase is variable. In this case of human cells, average cell cycle is of about 24 hours, mitosis takes

30 minutes, G1 phase 9 hours, S phase 10 hours, and G1 phase 4 hours and 30 minutes. In yeast the full cycle is of only 90

minutes.

Amitotic Cell Division in Prokaryotes

Prokaryotes (bacteria) reproduce by amitosis or binary fission (Fig No. 21.2). There is a single chromosomes

having a circular DNA molecule. When the DNA in bacterial cell is replicating, it is attached to the plasma membrane at two

separate points. Continued growth of the cell gradually separates the chromosomes, which are still attached lto the

membrane. Eventually, the plasma membrane and the bacterial cell wall grow inward, dividing the cell into two.

Cell Division

There are two types of cell division: Mitosis and Meiosis.

Mitosis in animal Cell

The word mitosis comes from the Greek word ‘mitos’ which means thread, and refers to the threadlike appearance

of chromosomes during this period. Mitosis was first studied by Walther Flemming in animals and by Strassburger in plants.

Definition: Mitosis can be define as “ The division of cell in such a manner that the chromosomes are duplicated and

distributed equally to the daughter cells”. The process of mitosis reproduces cells and distributes equal DNA to each

daughter cell. Thus, mitosis is a qualitative division of the cell. It takes place in the somatic (body) cells. The process of cell

division can be divided into two main phases, karyokinesis and cytokinesis.

Karyokinesis: Division of Nucleus.

Traditionally karyokinesis is studied in four stages, but actually it is a continuous process.

(1) Phophase (Gk: Pro; before)

(2) Metaphase (meta ; after)

(3) Anaphase (ana; again)

(4) Telophase (telo; end)

Intraphase: (Preparing the Scene)

Interphase (inter; between) is the phase between cell division. By G 2 stage the cell has doubled much of its cell

contents. The cytoplasm contains two microtubule organizing centre (MTOCs) each with a pair of centriole.

The chromatin resembles interoven fine threads. One or more nucleoli are present in the nucleus. Duplication of

chromosomes takes place DNA replicates. Synthesis of RNA and proteins occur. The chromosomes are mot visible under a

light microscope, as they are still in the from of lossely packed chromatin.

Prophase: Formatio of Mitotic Apparatus)

Activities in the nucleus: The chromosomes being to shorten and thicken, coiling upon themselves a process called

condensation. The condensation process continues throughout the prophase.

Nucleolus diappears: The synthesis of r-RNA ceases when that portion of the chromosome bearing rRNA genes, is

condensed, as a result the nucleolus diappears.

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Activities outside the nucleus: Early in the prophase, the two centriole pairs start to move a part, and continue to move until

they reach the opposite poles of the cell establisheing the bipolarity of the cell. Three sets of fibers i.e. microtubules

originate from each pair of centriole. The microtubules are composed of protein tubulin and RNA. Two sets of microtubules

(half spindle and spindle) compose the spindle. The polar microtubules do not interact with the chromosomes. They

interdigitate with polar molecules from opposite poles.

Nuclear membrane disappears: During the formation of the spindle the nuclear membrane breaks down, and its

components are reabsorbed into the endoplasmic recticulum.

Aster: When centrioles reach the poles of the cell, they radiate the third set of mictotubules outward, thus bracing the

centrioles against the cell membrane, this arrangement of microtubules is called aster. The function of the aster is brobably

mechanical (acting to stiffen the point of micro-tubular attachment during the later contraction of the spindle).

Mitotic apparatus: The aster, spindle and centrioles are collectively called the mitotic apparatus.

Chromatids: At the beginning of prophase the chromatin material is condensed and folded, as a result chromosomes appear

as thin threads which range in length from 0.25µm to 50 µm in length. By late prophase, each chromosome appears double.

The two halves are called chromatids. Each pair of chromatids have a centromere. Specialized protein complexes called

kinetochore having specific base arrangement develop on either side of each centromere. The second set of microtubules of

spindle fibres are called each centromere. The second set of microtubules of spindle fibres are called kinetochore spindle

fibres. The kinetochores of sister chromatids capture these fibers coming fron opposite poles. Forces associated with spindle

microtubules move the chromosomes toward centre of the cell.

Metaphase: (Division of Centromers)

The chromosomes move towards the equator of spindle, (half way between the poles, the equatorial or metaphase

plate). It is not a physical structure but an imaginary one. The kinetochore extends as spindle form fibers, the two poles of

the spindle are attached to the kinetochore of centromere.

The centromeres of all chromosomes are lined up on the metaphase plate. For each chromosome, the kinetochores

of the two sister chromatids face opposite pole of the spindle. The microtubules attached to a particular chromatid come

from one pole of the spindle, and those attached to its sister chromatid come from the opposite pole. At the end of the

metaphase, the centromeres divide, freeing the two sister chromatids from their attachment to one another. Centomere

replication is simultaneous for all the chromosomes.

Anaphase: (Separation of the Sister Chromatids)

The beginning of the Anaphsase is marked by the separation of the sister chromatids. Each sister chromatid now

rapidly moves towards the poles, to which its microtubule is attached.

The poles move apart: The spindle fiber consists of microtubules which occur in the form of a pair. Each member of the

pair is attached to opposite poles. The pair of microtubules of spindle fibre slide over one another, as a result the poles move

apart and increase in length.

The centromeres mover toward the poles: The kinetochore of the chromosomes are attached to the poles by half spindle

fibers, so with the movement of the pole apart, the chromosomes also move towards each pole. The spindle fibers do not get

shorten by condensation. At the end of the poles the spindle fiber is broken down into its subunits by the action of enzymes.

The tubulin from the spindle fibre is removed, as a result the spindle fiber becomes shorter and shorter, pulling the

chromosomes closer to the pole of the cell. Anaphase is over when equivalent and complete collection of chromosomes have

reached the two opposite of the cell.

Telophase: (Reformation of Nuclei)

At the beginning of the telophase the two sets of chromosomes reach the opposite poles of the cell. The condensed

and coiled chromosomes begin to uncoil. They increase in their length, and become thinner i.e. the chromosomes become

like the chromosomes of interphase stge.

Disappearance of spindle: The spindle fibers break into tubulin subunits reach the opposite poles of the cell. The

condensed and coiled chromosomes begin to uncoil. They increase in their length, and become thinner i.e. the chromosomes

become like the chromosomes of interphase stage.

Formation of the nuclear membrane: Each set of sister chromatids becomes suttounded by endoplasmic reticulum. This

ER form the nuclear membrane around each set of chromosomes.

Appearance of nucleolus: The chromosome in each set continue to uncoil. One of the early genes to regain expression are

the genes for rRNA. The rRNA are synthesized. These newly formed rRNA form the nucleoulus. At the end of the

telophase the nuclear division is over.

Cytokinesis: The Division of Cytoplasm

Nuclear division is over, but not the cell division. The new nuclei are still in the same cytoplasmic unit. Commonly

nuclear division is followed by separation of the cytoplasm into two parts. This separation accomplished by pinching of the

cell membrane when the astral microtubukles send signals to equatorial region of the cell, where actin and myosin are

activated which form the contractile ring. The pinching near the middle of an animal cell forms a cleavage furrow. The

process of cytoplasmic division is called cytokinesis. (Gk: Cyto; cell, Kinesis; movement). Cell organelles are distributed to

the towo daughter cells.

Significance of Mitosis

Diploid No: The diploid (2n) number of chromosomes is maintained in the cell.

Asexual reproduction: Asexual reproduction of single called eukaryotes takes place by mitosis.

Development: It is the basis of growth in the multicellular eukaryoptes.

Replacement of cells: Healing of wounds and replacement of damaged cells is accomplished by mitosis.

New cells: New cells are produced by mitosis.

Genetic continuity: The genetic informationi.e. DNA is regularly and equally distributed to the daughter cells.

Cancer: It is caused by uncontrolled mitosis.

TABL;E: 20.1 COMPARISON

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Mitosis in Animal Cell Mitosis in Plant Cell

Mitotic apparatus is Mitotic apparatus is

formed by aster, formed by spindle fibers

centriole and spindle only, as the aster and

fibers centrioles are absent in the

cells of higher plants.

In cytokinesis the It is achieved by the

cytoplasm is pinched in formatuion of a partition

the centre and forms two wall. The interzonal region

cells. spindle fibers at the

anaphase get transformed

into a structure called cell

plate or phragmoplast. It

extends round the equatorial

plate separating the two

cells.

Cancer

Cancer, is a disease of the cell cycle. Unlike normal cells of the body, cancer cells do not have properly functioning

cell-cycle control system and therefore divide excessively. This excessive growth can result in an abnormal mass of cell

called a tumor. Not all tumors are cancerous, however, a benign tumor is and abnormal mass of essential normal cells.

Benign tumors can cause problems if they grow in certain organs, Such as the brain but usually they can be completely

removed by surgery. They always remain at their original site in the body.

In contrast to a bengn tumor, a malignant tumor is cancerous. It is a mass of cancer cells, which are capable of

spreading into neighbouring tissues and often to other parts of the body. A malignant tumor arises from a single cancer cell

and displaces normal tissue as it grows. If the tumor is not killed or removed, some of the cancer cells spread into

surrounding tissue, enlarging the tumor. Cells may also split off from the tumor, invade the circulatory system (lymph

vessels and blood vessels), and travel to new locations, where they can form new tumors. The spread of cancer cells beyond

their original site is called metastasis.

Cancers are named according to the organ or tissue in which they arise, and over 200 different tupes are recognized

in humans. Form simplicity they are grouped into four categories. Carcinomas are cancers thet originate in the external or

internal coverings of the body; such as the skin or the lininig of intestine. Sarcoma arise in tissues that support the body,

such as bone and muscle. Cancers of blood-forming tissues, such as bone marrow, spleen are called leukemias and that of

lymph nodes are called lymphomas. Cancer cells can go on dividing indefinitely, as long as they have a supply of nutrients.

If cancer cells stop dividing, the seem to do so at random points in the cell ctycle, rather than just at the normal cell-cycle

checkpoints.

Treatment

Two types of cancer treatment, chemotherapy and radiation therapy, attempt to halt the spread of cancer cells by

stopping them from dividing. In radiation therapy, parts of the body that have cancerous tumors are exposed at high-energy

radiation, which disrupts cell division. Chemotherapy uses the same strategy as radiation; in theis case, drugs that disrupt

cell division are administered to the patient.

Cancer cells can be distinguished from normal cells because they have high nucleus to cytoplasm ratio, prominent

mnucleoli and many mitosis. The presence of invading cells in otherwise normal tissue is an indication of malignancy.

Cancer is caused mainly mutations in somatic cells. Secondary, the cancer results from the accumulation of a few as three to

as many as twenty mutations, in genes that regulate cell division.

Characteristic of Cancer Cells

At the least four features characterize all cancer cells.

(1) Their plasma membrane and cytoplasm change profoundly.

(2) Cancer cells grow and divide abnormally.

(3) Cancer cells have a wakened capacityu for adhesion.

(4) Cancer cells are lethal.

Meiosis in an animal cells

Sexual reproduction takes place in animal and plants by forminggametes. The gametes fuse to form zygote. They

zygote forms the organism. The number of chromosomes remain constant in the successive generations. How the number of

chromolsomes remoan constant from generations to generations? August Weismann proposed the hypothesis that “ there

must be a kink of cell division in which the chromosome member is halved”. There are two types of cell. (i) Somatic cells

which make up the body. (ii) Germline cells which produce the gametes. Both germline cells and somatic cells are diploid

(2n). When a germline cell undergoues division it produces cells with half of the diploid number of chromosomes i.e.

haploid (n) and the process of division is called reduction division or meiosis. Sperms and eggs are produced by meiosis in

anuimals. Spores are produced by meiosis in plants.

Definition: Meiosis is a process of nuclear division in which the number of chromosomes in a cell is halved during cell

division.

The Stages of Meiosis

Meiosis is a continuous process. It can be described most easily by dividing it into two arbitrary stages. The two

stages of meiosis are called meiosis I and meiosis II. These are further subdivided into prophase, metaphase, anaphase and

telophase.

The First Meiotic Division

Interphase I: The DNA duplicates by its replicating process. The fine chromosomes. It is further divided into five stages:

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(1) Leptoten (2) Zygotene (3) Pachytene (4) Dipotene (5) Diakinesis.

Leptoten: (Thin Thread Stage)

Chromosomes look like interweaving threads. Due to condensation of chromatin material; chromosomes become

more apparent and distinct, nucleus increases in size. Initially thin chromosomes become shorter and thicker. The

chromosomes in size. Initialy thin chromosomes become shorter and thicker. The chromosome number of the cells is seen.

Homologous chromosomes start getting closer to each other. In the cell there are two of each type of chromosome. The

identical chromosomes are called homologous chromosomes (Gk: Homologue; agreement). The morphology and position

of the centromere of the homologous chromosome is same. For example in man, the number of chromosome is 46 and the

homologous pair of chromosome is 23.

Zygotene: (Pairing Stage)

The homologous chromosomes begin to pair length wise with its homologue. The process of pairing is called

synapsis. (si-nap-sis) (Gk: Union). The synapsis may start from any point with the corresponding complementary DNA stand

of the other.

Pachytene: (Thickening Stage)

After the completion of synapsis the chromosomes shorten and thicken due to condensation of the chromatin

material land their double nature is evident. Each pair of synapsed chromosome consists of four chromatids, two

centromeres and is called a tetrad or bivalent.

Crossing over: The chromatids of the homologues my cross each other and the point of crossing is X shaped figure under

the light microscope. It is called chiasma (Gk: chiasma; cross, plural: chiasmata). Chromosomes segment is exchanged

between the two homologous chromosomes at the chiasma and is called crossing over.

Diplotene: (Duplication Stage)

The paired homologous chromosomes begin to separate by repelling. (The bivalent is still attached at chiasmata or

chiasma). Diplotene can last for months or years. The chromatids are clearly visible due to the progress in condensation

cycle.

Diakinesis: (Moving Apart Stage)

The tetrad are more evenly distributed in the nucleus against the nucleoli envelope. The number of chiasmata

begins ro reduce as the diakinesis reaches to its end, but the terminal chiasmata are still present. The nucleoli disappear. The

nuclear membrane is still present. The diad (the two chromatids attached to a single centromere is called diad) behaves as

a single unit because they are held together by a common centromere, throughout meiosis.

Metaphase I

The nuclear membrane disintegrate. The microtubules form the spindle. The chromosome lineup in double row

i.e. pair on the equator. Microtubules bind only to one kinetochore of each centromere. The centromere of one homologue

becomes attached to microtubules extending to one pole, whereas the centromere of the other homologue becomes attached

to microtubules extending to other pole.

Anaphase I

The attachmend of chromosomes to the spindle is complete. The microtubules that are attached to the

homologous chromosomes begin to slide over one another, as a result the spindle poles move apart. The spindle fibers

become shorter as the spindle fibers are dragged to the poles. They are broken by the action of enzymes. With the

shortening of spindle fibers, the chromosomes that are attached to the spindles are also pulled towards each pole. When the

shortening of spindle fiber is completed, each pole has single set of chromosomes, consisting of one member of each

homologous pairs.

Telophase I

In telophase I, each set of chromosomes is present at their respective poles. By cytokinesis two cells are formed,

which are haploid.

The second Metotic Division: Meiosis II is simply a mitotic division. It occurs to separate the sister chromatids as in

mitosis.

InterphaseII: DNA does not duplicate in interphase II interphase II is very brief. Each of the two cells resulting from

meiosis I progress into meiosis II very quickly.

ProphaseII: The complicated nuclear events of prophase do not take place.

Metaphse II and Anaphase II: The chromosomes lineup in the same fasion as they did in metaphase I. Here again a randon

distribution of chromosomes take place. The centromere divides, as a result the two chromatids are seoarated. As each

chromatid is now a sparated structure they are called chromosomes. Chromosomes are distributed in equal number, to each

pole. DNA does not replicate but the centomere divides.

Telophase II: The nuclei are reconstructed in the typical manner. Each nucleus now constains haploid set of chromosomes,

because DNA has duplicated only once during the cell division.

Fate of haploid cells: Each of the four haploid products of meiosis contains a basic set of chromosomes. These haploid cells

may function directly as gametes, as they do in animals or may continue to divide by mitosis, as they do in plants, fungi and

many protests.

Table: 20.2 Comparison

Mitosis Meiosis

1. It takes place in somatic 1. It takes place in the

cells. germline cells

2. It consists of one division 2. It consists of two divisions

3. The centromere divides in 3. The centromere does not

mitosis at metaphase divide at metaphase of

meiosis I.

4. The chromosomes are 4. The chromosomes are

monovalent in the bivalent in the daughter

dauthter cells cells of meiosis I.

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5. The number of 5. The number of

chromosomes remains chromosomes is halved in

constant meiosis I and this half

number remains same in

meiosis II.

6. Chiasmata, tetread, 6. Chiasmata, tetrad, crossing

crossing over do not take over take place

place

7. Two diploid cells are 7. Four haploid cells are

formed formed

8. The daughter cells remain 8. The daughter cells are

as cells transformed into sperms or

eggs cells.

9. Genetic content of mitotic 9. Genetic content of meiotic

products is identical products is different

10. Mitotic products are 10. Meiotic products cannot

usually capable of underfo further meiotic

undergoing additional divisions.

mitotic divisions

Significance of Meiosis

Reduction of chromosome number:

The number of chromosomes become half (haploid) in the gametes due to which the number of chromosomes

remain constant from generation to generation.

Distribution of genetic material: It provides a mechanism for the transmisision of genetic material form one generation to

the next.

Mixing of genetic material: There is a mixing of genetic material in the zygote formed by the fusion of gametes from

different parents. As a result there is also genetic variability.

Role in adaptation: It is the meiosis which offers a chance for the production of the best adapted individual and better

chance for survival and better chance for survival in the changed environment.

Role in evolution: Meiosis provides a raw material for evolution. Evolution is impossible without a change. Meiosis serves

as a source for the change.

Non-Disjunction

Meiosis occurs repeatedly, as testes or ovaries produce gametes. Almost always, the meiotic spindle distributes

chromosomes to daughter cells without an error. But occasionally there is an accident, called a nondisjuction, in which the

members of a chromosome pair fail to separate. Figure No. 21.9. A and B illustrate two ways that nondisjuction can occur.

For simplicity, we use a hypothetical organism whose diploid chromosome number is 4. In both figures, the cell at the top is

diploid (2n), with two pairs of homologous chromosomes undergoing meiosis I.

Sometimes, as in Figure 21.9, a pair of homologous chromosomes does not separate during meiosis I. In this case,

even though the rest of meiosis occurs normally, all the resulting gametes end up with abnormal numbers of chromosomes.

Two of the gametes have three chromosomes, two of which are the same; the others two gametes have only one

lchromosome each. In figure B, meiosis I is normal, but a pair of sister chromatids fail to move apart in one of the pair of the

cells during meiosis II. In this case, two gametes have the normal complement of two chromosomes each, but the other two

gametes have the abnormal complement.

Figure 21.9 shows what happens when an abnormal gamete produced by nondisjuction unites with a normal gamete

in fertilization. Here, an egg cell with two copies of one of its chromosomes (a total of n + 1 chromosomes) is fertilized by a

normal sperm cell (n). The resulting zygote has an extra chromosome (a total of 2n + 1 chromosomes). Mitosis will then

transmit the abnormality to all wmbryonic cells. If this were a real organism and if survived, it would have an abnormal

karyotype and probably a syndrome of disorders caused by the abnormal number of genes. Nondisjunction can lead to an

abnormal chromosome number in either sex of any sexually reproducing, diploid organism, including humans. Three

chromosome disorders of interest are Down syndrome, Klinefelter syndrome and Turner syndrome, Klinefelter syndrome

and Turner syndrome.

Down Syndrome (Trisomy 21)

Down syndrome (Down’s is not used these days) is also known as Mongolism (Fig: 20.10) is easily recognized by

these characteristics: short stature; an eyelid fold; stubby fingers; a wide gap between the first and second toes; a large,

fissured tongue; a round head; a palm crease, the so-called simian line; and, unfortunately, mental retardation, along with an

enlarge tongue. Down syndrome is also called trisomy 21 because the individual usually has three copies of chromosome 21.

In most instances, the egg had two copies instead of one of this chromosome. The chances of a woman having a Down

syndrome child increase rapidly with age, starting at about age 35. The frequency of Down syndrome is 1 in 800 births for

mothers under 40 years of age and 1 in 80 for mothers over 40 years of age.

Turner Syndrome (XO)

Turner Syndrome occurs in one in 6,000 births. The individual is XO with one sex chromosome, in XO the O

signifies the absence of a second sex chromosome. These females are short, have a broad chest, and webbed neck. The

ovaries, oviducts, and uterus are very small and nonfunctional. Turner females do not undergo puberty or menstruation, and

there is a lack of breast development. They are usually of normal intelligence and can lead fairly normal lives, but they are

infertile even if they receive hormone supplements.

Klinefelter Syndrome (XXY)

Klinefelter syndrome occurs in one in 1,500 briths. These males with two or more X chromosomes in addition to a

Y chromosome are sterile. The testes and prostate gland are underdeveloped, and there is no facial hair. Also, they may be

some breast development. Afffected individuals have large hands and feet and very long arms and legs. They are usually

slow to learn but mentally retarded unless they inherit more than tow X chromosomes.

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Apoptosis and Necrosis

The first cell of a new multicelled individual contains marching orders that will guid its descndants along a program

of growth, development, and reproduction, then on to death. As part of the program, many cells proceed to self-destruction

when they compete a prescribed function. If they become altered in ways tat might pose a threat to the body as a whole, as

by infection or cancer, they can execute themselves. Apoptosis (Gk; dropping off or falling off) is the name for this form of

cell death. It starts with molecular signals that activate and unleash lethal weapon of self extraction, which were stockild

earlier within the cell. Protein cleavage enzymes are such weapons. Think of them as folded pocketknives. When popped

open, they chop apart structural proteins, including the building blocks of cytoskeleton elements and nucleosomes that

organize the DNA.

The dying cells shrink, condense and ultimately split up, thus release small membrane bounded apoptic bodies,

such are generally phagocytosed by other cells. Intracellular constituents are not released freely in extracellular atmosphere

which otherwise might have deleterious effects. In contrast to suicide the cell death due to tissue damage is called necrosis

during which the typical cells swells and bursts, releasing the intracellular contents, which can damage neighbouring cells

and cause inflammation.

EXERCISE – SECTION – I – OBJECTIVE QUESTIONS

1. Fill in the blanks

i. During cell division. ____ sets of microtubules originate from each pair of centrioles.

ii. In Down syndrome, chromosme no. ____ fails segregate.

iii. A person with Klinefelters syndrome has an extra _____ chromosome.

iv. The cell death due to tissue damage is called ______.

v. Division of the nucleus is called ______.

vi. Mitotic apparatus is formed during ______.

vii. Spindles are composed of mainly a protein called ____.

viii. In man there are ____ pair of homologous chromosomes.

ix. Exchange of chromosomal segment is called ___.

x. In plants meiosis takes place during the formation of ____.

2. Mark the statements as “True” or “False”.

i. Interphase is the period of great biochemical activity.

ii. Meiosis occurs in haploid cells.

iii. During S-phase, DNA is synthesized and chromosome number is doubled.

iv. Division of the cytoplasm is called cytokinesis.

v. Equatorial plate is formed in plants.

3. Select the correct answer and encircle it.

i. It is period of extensive metabolic activity: (a) G0 (b) G1 (c) G2 (d) S

ii. The period of cell cycle between two consecutive divisions is termed as: (a) prophase (b) metaphase (c) telophase (d)

interphase

iii. Cancer is caused mainly by mutations in: (a) somatic cells (b) sex cell (c) G 2 Stage (d) S-phase

iv. The sex chromosme complement in individuals with klinefelter syndrome is: (a) XXY (b) XY (c) XXYY (d) XYYY

v. Mitosis is divided into: (a) karyokinesis (b) cytokinesis (c) interphase (d) both a and b

vi. Small localized tumors are called: (a) benign (b) malignin (c) cancer (d) interdigitate

vii. The significane of mitosis is: (a) to bring about cell division (b) to produce daughter cells with the same genetic make

up as the parent cell. (c) to control the size of the cell (d) to control cell contents.

viii. Which of the following is not for crossing over? (a) it occurs during prophase I (b) it occurs during prophase II (c) it

occurs between homologues (d) it is seen as X-spahped structure called chiasmata.

4. Match the following columns.

Column ‘A’ Column ‘B’

a. Phragmoplats i. Somatic cells

b. Cancer ii. Panchytene

c. Tetrad iii. Anaphase I

d. Crossing over iv. Plant cell

v. Zygotene

5. Draw a label:

(i) stages of mitosis. (ii) stages of meiosis

SECTION – II – SHORT QUESTIONS

(1) Distinguish between:

mitosis and meiosis - chromosme and chromatin

chromosome and chrmatids – centromere and centriole

cytokinesis and karyokinesis – centromerere and kinetochore

chiasmata and chiasma – diad and bivalent

apoptosis and necrosis – haploid and diploid number of chromosomes.

2. Write notes on:

karyotypes chromatine kinetochore

cytokinesis crossing over leptotene

zygotene pachytene diplotene

diakinesis apoptosis Down syndrome

klinefelter syndrome turner syndrome cancer

nondisjuctionof chromosme

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3. Write the differences between:

(a) mitosis in animal cell and plant cell

(b) Mitosis and meiosis

4. Write the significance of (a) mitosis (b) meiosis

5. What is mitotic apparatus?

6. How does meiosis contribute to genetic recombination?

7. What is synapis and crossing over?

8. Define cell cycle and outline main features of each stage.

SECTION – III – EXTENSIVE QUESTIONS

1. Give a detail account of mitosis in an animal cell.

2. Give a detail account of meiosis in an animal cell.

3. Write a comprehensive note on cancer.

4. Give a detailed account of nondisjunction, and the syndromes caused.

ANSWERS

1.

(i) 3 (ii) 21 (iii) X

(iv) necrosis (v) karypkinesis (vi) prophase

(vii) tubulin (viii) 23 (ix) crossing over

(x) spores

2.

(i)T (ii) F (iii) T (iv) T (v) F

3.

(i) b (ii) d (iii) a (iv) a (v) d (vi) a

(vii) b (viii) b

4.

(a) iv (b) I (c) v (d) ii



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