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1 Genetics of Forensic DNA Typing

2 2-12 September 1997

3 Course Description

4

5 Course Description/Prerequisites:

6 This is for forensic DNA practitioners, those interested in a career in forensic DNA and all stude

7 interested in broadening their understanding of the genetic principles that underlie the forensic application

8 DNA typing methods. Attendance by by crime laboratory staff and other interested biology and law stude

9 and professionals is expected.

10 This two week (72 hour) course is a combined lecture and laboratory (demo) course designed to prov

11 comprehensive exposure to the fundamentals of classical, molecular and population genetics with applicatio

12 specifically for practicing forensic DNA analysts. It will include the principles underlying methods used

13 isolate, quantitate and analyze DNA from biological materials (including RFLP and PCR) that enable

14 detection and analysis of polymorphic regions. The course is divided into the following sections: I. Classi

15 Genetics to include mendel's principles, meiosis and mitosis, sex determination, sex linkage, chromosom

16 mapping and cytogenetics, II. Molecular Genetics to include structure and chemistry of DNA, methods used

17 study DNA (gel and capillary electrophoresis, restriction enzymes, reassociation kinetics, southern b

18 analysis, cloning, PCR, sequencing, and SBH) transcription, translation, developmental genetics/control of ge

19 expression, DNA mutation and repair, recombination, transposable elements, extrachromocomal (n

20 mendelian) inheritance (mitochondrial DNA), and applications of recombinant DNA with an emphasis

21 forensic DNA typing including introductions to the following forensic genetic marker systems: VNT

22 PM+DQA1, D1S80, and STR typing and III. Population Genetics to include measuring, genetic variati

23 quantitative genetics, inbreeding, mutation, genetic drift, gene flow, Hardy-Weinberg Equilibrium, foren

24 DNA statistics, molecular evolution, and conservation genetics.

25 Critical thinking will be emphasized through problem solving, student assignments and gro

26 discussions. Demonstrations will highlight methods currently being utilized in forensic DNA laborator

27 One exam will be given and the results of the exams will be available after the course.

28

29 Each class is limited to 40 students.

30 This course is POST reimbursable , PLAN IV (to be determined)

31

32 Teaching Methods: Classroom lectures, group discussions, laboratory exercises and exams

33 Student Objectives: Students will be required to demonstrate their knowledge in written

34 assignments, group problem solving and exams.

35 Grading: Letter grades based on final exam and assignments. Attendance at all

36 sessions is mandatory.

37 Prerequisites: A class in molecular biology or biochemistry is highly recommended

38 Preparation: Pre-course reading material. RF. Weaver and PW. Hedrick. 1997.

39 Genetics. Third Edition. Wm. C. Brown Publishers.638 pages. ISBN 0- 697-

40 16000-9

41 Location: Lectures: Trailer I Temp. Class 1, UC Davis Campus Davis

42 Laboratories: 143 Briggs Hall

43

1 Genetics of Forensic DNA Typing

2 2-12 September 1997

3 Course Outline

4

5 2 September 1997

6 8:30 WELCOME Steve Lee

7 Introduction of Participants

8 Sign In

9 Overview of Course

10

11 8:45 LECTURE Introduction to Genetics Steve Lee

12 Transmission Genetics 2-15, 73-87

13 Basic Terminology

14 Chromosome structure

15 Cell Cycle

16 Molecular Genetics

17 Discovery of DNA

18 What genes are made of and do

19 Relationship of genes and proteins

20

21 10:30 BREAK

22

23 10:45 LECTURE Introduction to Genetics - Continued

24 Mitosis, Meiosis

25

26 12:00 LUNCH

27

28 13:00 LECTURE Classical Genetics - Mendel's Principles Steve Lee

29 Segregation 18-27

30 Independant Assortment

31 Punnett Squares

32 13:45 BREAK

33

34 14:00 LECTURE Introduction to Probability and Statistics Keith Inman

35 Basic vocabulary of probability 28-38

36 Product rule

37 Sum rule

38 Basic statistics and hypothesis testing

39

40 14:45 BREAK

41

42 15:00 LECTURE Statistics Problem solving Keith Inman

43

44 15:45 DISCUSSION Questions- Student Assignments

45

46 16:30 ADJOURN

47

1 Genetics of Forensic DNA Typing

2 2-12 September 1997

3 Course Outline

4

5 3 September 1997

6 8:30 REVIEW Lee/Inman

7 Review of Terminology, Meiosis, Mitosis

8 Review of Mendel's principles

9 Review of Basic Probability/Statistics

10 Questions/Discussion

11 Review Student assignments

12

13 9:30 LECTURE Classical Genetics - Mendel's Principles Steve Lee

14 Sex chromosomes 100-101

15 Rules of Expression 42-50

16 Polygenic inheritance and environmental effects 50-69

17 10:15 BREAK

18

19 10:30 LECTURE Applications of Mendel's Principles Steve Lee

20 Dominance, Incomplete, Codominance 42-50

21 Lethals

22 Pleiotropy

23 Penetrance and Expressivity

24 Multiple alleles

25 12:00 LUNCH

26

27 13:00 LECTURE Application of Mendel's Principles Cont Lee

28 Sex Linked Genes 50-69

29 Epistasis

30 Genotype-Phenotype

31 Mean/Variance Quantitative Traits

32 Estimating Genetic Variance

33 Paternity

34

35 14:15 BREAK

36 14:30 LECTURE Genetic Linkage and Recombination Lee

37 A physical explanation 104-125

38 Rate of Recombination

39 Linkage Map

40 Tetrad analysis

41 Crossing Over

42 15:45 BREAK

43

44 16:00 DISCUSSION Questions- Student Assignments Lee

45 16:30 ADJOURN

46

1

2 Genetics of Forensic DNA Typing

3 2-12 September 1997

4 Course Outline

5

6 4 September 1997

7 8:30 REVIEW Steve Lee

8 Review of Sex Chromosomes

9 Review of Applications of Mendel's principles

10 Review of Linkage and Recombination

11 Questions/Discussion

12 Review Student assignments

13

14 9:30 LECTURE Molecular Genetics Steve Lee

15 DNA Structure 130-150

16 Introduction to gene function

17 Basic Mechanism of DNA Synthesis

18 Semiconservative

19

20 10:30 BREAK

21 10:45 LECTURE Molecular Genetics Continued Lee

22 Enzymology of DNA Replication 154-177

23 Recombination of Genes 177-183

24 12:00 LUNCH

25

26 13:00 LECTURE Molecular Genetics Continued Lee

27 Replication and Recombination 177-183

28 13:45 BREAK

29

30 14:00 LECTURE Transcription and Control in Prokaryotes Lee

31 Fundamentals 185-215

32 RNA Polymerases and Promoters

33 Operons

34 Control of transcription

35 DNA-protein interactions

36 14:45 BREAK

37 15:00 LECTURE Application of Gene Cloning (and PCR) Pamela Ronald

38 Agriculture Ch 20

39 Molecular analysis of

40 Pathogen/Plant interactions

41

42 16:00 DISCUSSION Questions- Student Assignments Lee

43 16:30 ADJOURN

44

1 Genetics of Forensic DNA Typing

2 2-12 September 1997

3 Course Outline

4

5 5 September 1997

6 8:30 REVIEW Steve Lee

7 Review of DNA Structure/Function/Synthesis

8 Review of Prokaryotic Transcription

9 Reviw of Methods used to Study DNA

10 Questions/Discussion

11 Review Student assignments

12

13 9:30 LECTURE Molecular Genetics John Tonkyn

14 Eukaryotic Gene Structure 220-234

15 Chromatin Structure

16 Nucleosomes

17 RNA Polymerases and their roles

18 Regulation of Transcription

19 1045 BREAK

20 11:00 LECTURE Molecular Genetics cont John Tonkyn

21

22 Gene-specific factors for RNA Pol II 234-252

23 Relationship btwn Structure:Expression

24 Post-transcriptional Modifications

25 12:00 LUNCH

26

27 13:00 LECTURE Molecular Genetics- Translation Lee

28 Protein Structure/Function 255-271

29 Ribosomes and ribosomal proteins

30 tRNA 2o and 3o structure of tRNA

31 Amino acid binding to tRNA

32 Amino acyl tRNA synthetases

33 14:00 LECTURE Molecular Genetics- Translation continued Lee

34 The Genetic Code-Triplet code 271-292

35 Breaking the Code- The (Almost)

36 Universal Code

37 Mechanism of Translation

38 Translational Control

39 15:00 BREAK

40 15:15 LECTURE Methods Used to Study DNA Inman

41 DNA Extraction

42 Agarose Gel Electrophoresis 146-149

43 15:30 LABORATORY Demonstration of DNA Extraction and

44 Agarose Gel Electrophoresis Inman/Brewer/

45 Renfroe/Speer

46 17:30 ADJOURN

47

1 Genetics of Forensic DNA Typing

2 2-12 September 1997

3 Course Outline

4

5 8 September 1997

6 8:30 REVIEW Steve Lee

7 Review of Eukaryotic Transcription

8 Translation

9 Review of DNA Extraction/Gel Electrophoresis

10 Questions/Discussion

11 Review Student assignments

12

13 9:30 LECTURE Molecular Genetics- Gene Mutation Lee

14 Types of Mutations 295-299

15 How Mutations Affect the Genetic Material 299-305

16 Triplet repeats 305-306

17 Mutagenesis306-309

18 Silent Mutations/Reversions 309-310

19 1045 BREAK

20

21 11:00 LECTURE Molecular Genetics- Gene Mutation Lee

22 DNA Repair 311-318

23

24 12:00 LUNCH

25

26 13:00 LECTURE Methods Used to Study DNA Orrego

27 DNA-DNA Reassociation Kinetics 142-147

28 Restriction Fragment Length Polymorphisms (RFLPs)

29 Cloning 123-125

30 Southern/Northern/Western Analysis 415-441

31 DNA Sequencing

32 Polymerase Chain Reaction (PCR) 459-460

33

34 14:15 BREAK

35

36 14:30 LECTURE Methods Used to Study DNA Orrego

37 Quantitative aspects of PCR

38 Direct Sequencing

39 Single Stranded Conformational Polymorphisms

40 Denaturing Gradient Gel Electrophoresis

41 15:00 BREAK

42

43 15:15 LABORATORY Demo - Setting up PCR Inman/Renfroe

44 of Short Tandem Repeats /Speer/Brewer

45 16:00 DISCUSSION Questions- Student Assignments

46 16:30 ADJOURN

47

1 Genetics of Forensic DNA Typing

2 2-12 September 1997

3 Course Outline

4

5 9 September 1997

6 8:30 REVIEW Steve Lee

7 Review of Gene Mutation

8 Review of Methods used to Study DNA

9 Review of PCR

10 Questions/Discussion

11 Review Student assignments

12

13 9:30 LECTURE Molecular Genetics - Applications of DNA Gary Sims

14 in forensics: Casework - People vs OJ Simpson

15 An overview of the Forensic Uses of 475-478

16 DNATyping

17 VNTRs 459

18 11:15 BREAK

19 11:30 LECTURE Applications of DNA in forensics (cont) Gary Sims

20 Reverse Dot Blot typing systems

21

22 12:00 LUNCH

23 13:00 LECTURE Introduction to D1S80 Pardo/Staff

24 History

25 Advantages in Forensics

26 Discrimination

27 Detection Methods

28 13:45 BREAK

29 14:00 LECTURE Introduction to Short Tandem Repeats Lee/Tonkyn

30 Polymorphisms 460

31 Loci-Multiplexes

32 Applications in Medicine

33 Advantages in Forensics

34 Discrimination

35 Detection Methods

36 Polyacrylamide Gel and Capillary Electrophoresis

37 14:45 LECTURE Quality Control, Quality Assurance, Lee

38 Accreditation of DNA in Forensics

39 Validation, TWGDAM, DNA Advisory Board

40 American Society of Crime Laboratory Directors

41 15:15 LABORATORY Demo: Setting up RFLP of PCR Products Inman/Renfroe

42 16:00 DISCUSSION Questions- Student Assignments /Speer/Brewer

43 16:30 ADJOURN

44

1 Genetics of Forensic DNA Typing

2 2-12 September 1997

3 Course Outline

4

5 10 September 1997

6 8:30 REVIEW Steve Lee

7 Review of Application of DNA in Forensics

8 Review of PCR based DNA Typing Systems

9 Review of QC, QA in Forensics

10 Questions/Discussion

11 Review Student assignments

12

13 9:30 LECTURE Molecular Genetics-Transposons Steve Lee

14 Bacterial Transposons 322-332

15 Insertion Sequences

16 Eukaryotic Transposons 332-337

17 1045 BREAK

18

19 11:00 LECTURE Molecular Genetics- Extrachromasomal

20 mtDNA Inheritance Ellen Clark

21 Cytoplamic Inheritance 504-518

22 Mapping of mitochondria and

23 chloroplasts

24

25 12:00 LUNCH

26

27 13:00 LECTURE Molecular Genetics- Extrachromasomal Ellen Clark

28 Molecular Genetics of Organelles 518-526

29 Origin of Mitochondria and Chloroplasts 526-529

30 14:15 BREAK

31

32 14:30 LECTURE Application of Gene Cloning (and PCR) Lee

33 Human Disease Genes- PCR used in diagnostics

34 Advanced Genetic Mapping 448-475

35 15:15 BREAK

36

37 15:30 LECTURE Application of Gene Cloning (and PCR) Lee

38 Pharmaceuticals/ Medicine

39 Human Genome 457

40

41 16:15 DISCUSSION Questions- Student Assignments

42 16:30 ADJOURN

43

1 Genetics of Forensic DNA Typing

2 2-12 September 1997

3 Course Outline

4

5 11 September 1997

6 9:00 REVIEW Steve Lee

7 Review of Transposons

8 Review of Extrachromasomal Inheritance

9 Review of Applications of Gene Cloning and PCR

10 Questions/Discussion

11 Review Student assignments

12

13 10:00 LABORATORY LABORATORY OF STUDENT MATING Lee

14 1. To understand the following concepts: allele

15 frequencies; genotype frequencies; gene pool;

16 genetic drift; Hardy-Weinberg Equilibrium

17 2. To participate in a simulation of the evolutionary

18 process

19 3. To observe the effect of lethal genes, advantageous

20 genes, and small populations on allele frequencies

21 in a population

22

23 12:00 LUNCH

24

25 13:00 LECTURE Population Genetics Sensabaugh

26 Types of Genetic Variation 532-545

27 Measuring Genetic Variation

28 Hardy Weinberg Equilibrium

29 Inbreeding

30 Mutation

31

32 14:30 BREAK

33

34 14:45 LECTURE Population Genetics (Cont) Sensabaugh

35 Genetic Drift 545-554

36 Gene Flow

37 Natural Selection

38

39 15:45 LECTURE Forensic DNA Statistics Sensabaugh

40 17:00 DISCUSSION Questions- Student Assignments 557-558

41 17:30 Adjourn

42

43

1 Genetics of Forensic DNA Typing

2 2-12 September 1997

3 Course Outline

4

5 12 September 1997

6 8:30 REVIEW Steve Lee

7 Review of Pop Gen- Genetic Variation

8 Review of Hardy Weinberg

9 Review of Inbreeding, Mutation, Genetic Drift,

10 Gene Flow, Natural Selection

11 Review of Application of Population Genetics

12 Questions/Discussion

13 Review Student assignments

14

15 10:00 Extensions and Applications of Population Genetics Orrego

16 Frequency of Human Genetic Disease 558-577

17 Molecular Evolution

18 Conservation Genetics

19

20 11:00 Break

21

22 11:15 Questions/Student Review of topics Lee

23

24 12:00 LUNCH

25

26 13:00 EXAM Final Exam Lee/Staff

27

28 15:00 BREAK

29 15:30 "LABORATORY" Detection/Analysis of STRs

30 Interpretation Issues

31 Stutter/Repeat Slippage Lee

32

33 16:30 COURSE EVALUATIONS Lee

34

35 17:00 ADJOURN

36

37


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