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Bio102: Introduction to Cell Biology and Genetics

Genetic Code and Mutation



Key Terms:

inborn error of One Gene-One Start and Stop mutation mutagen

metabolism Enzyme Codons

metabolic pathway wildtype (wt) codon missense mutation carcinogen

minimal media transcription redundancy nonsense mutation Ames test

auxotroph translation reading frame frameshift mutation reversion

Central Dogma genetic code mRNA silent mutation



Key Figures: Figures 17.2, 17.3, 17.4, 17.5, 17.23

Lecture Outline:

Archibald Garrod describes “inborn errors of metabolism”

blocks a metabolic pathway, causing accumulation of an intermediate in the pathway

Beadle and Tatum experiment

induced mutations in Neurospora crassa with x-rays

found mutants that were arginine (an amino acid) auxotrophs, so they needed added arg to live

some could live with ornithine or citrulline, precursors in the arginine metabolic pathway

deduced that a gene corresponds to one enzyme, thus the One Gene-One Enzyme Hypothesis



Central Dogma: information moves from DNA (gene)  RNA  Protein



Genetic Code describes which three-nucleotide codon on the mRNA corresponds to which amino acid

code is universal and redundant (so one amino acid may be encoded by >1 codon)

always starts with AUG codon (=Met amino acid)

each codon is then translated in-frame

ends with either UAA, UGA or UAG (which do NOT correspond to an amino acid)



Mutation is a change in the DNA. May effect the protein sequence.

Missense mutations change one

amino acid (e.g. UAC  CAC)

Nonsense mutations change an

amino acid codon to a stop

codon (e.g. UAC  UAA)

Insertions or deletions cause a

change in the reading frame

(e.g. UAC  UA)

Silent mutations make no change in

the protein (e.g. UAC  UAU)





Ames test is used to find mutagenic

chemicals.



Measures mutation rate of a

reversion that allows Salmonella bacteria to no longer by a histidine auxotroph.



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