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From Wikipedia, the free encyclopedia Mate Choice









Mate Choice

Mate choice is the process in which one organism choos- 78% of their energy daily towards mate choice. Results

es another one to mate with. The overall reason for mate also suggest that assessing high quality mates are more

choice in any species is to pick a mate that can provide costly than assessing lower quality mates. This is due to

better genetic material that will produce more fit off- more time being taken to visit the higher quality mates.

spring that are more likely to survive and have offspring Not only was time lost, but also more body mass was

of their own. Mate Choice is based on the Theory of Nat- lost. In contrast, females that visited these higher quality

ural Selection. Only organisms that are well adapted to males showed a strong trend toward producing follicles

survive in the environment that they live in will survive of smaller size, which is something directly related to off-

long enough to mate and produce their own offspring, to spring survival. Even though a large percentage of ener-

pass along their traits. gy was used to choose a mate, the females did end up hav-

ing more fit offspring.

Direct and indirect benefits

Sexy Sons Hypothesis

Direct Benefits The Sexy Sons Hypothesis suggests that females obtain

Direct benefits of choosing mates affect the female di- future fitness benefits from mating with males through

rectly, such as increasing her own lifetime reproductive the inheritance of her son’s of traits.[8] Because genes are

output.[1] Females that choose males that are more able passed down from generation to generation, the proba-

to reproduce with them, then in turn increase the proba- bility of offspring to have a trait would increase greatly

bility of having more offspring that will survive.[2] if one parent or both parents had the trait to pass down.

Significant sire effects on egg to adult viability and son’s

Indirect Benefits pre-productive survival, as well as moderate to high coef-

Indirect benefits of mate choice are ones that benefit off- ficients of additive genetic variation for these traits have

spring of the organism making the choice. The organ- been found in Dung Beetles.[9] Beetles who have select-

ism chooses a mate with a particular favorable trait. This ed mates with better genetics tend to have offspring that

trait is then passed down to its offspring, benefiting the survive longer and are more able to reproduce than those

chooser directly. In some fish species, males with eggs that do not pick mates with genetic quality. This suggests

already in their nests were preferred over other males that carefully choosing a mate is beneficial.[10]

who had none. The indirect benefit that can be seen from

this example is that more offspring will survive and be

able to reproduce because the father is better at watching Types of Choice

over the eggs.[3] Choosing males that already watched

over eggs benefitted the mother indirectly because her

offspring now had a better chance to survive. Cryptic Choice

While most mate choice involves choosing of mates be-

Costs fore copulation has commenced, cryptic choice involved

selection after the organism has mated. Variation in rel-

Mate choice while it has its benefits, can be a costly un- ative male fertilization success is an important source

dertaking involving considerable expenditure of time in variance of male reproductive success.[11] The most

and energy.[4] This not only occurs in female mate prominent method of cryptic female choice is sperm

choice, but in male mate choice as well.[5] Optimal mate choice, when one male’s sperm is chosen over another

choice involves a tradeoff between the cost of selecting a male’s sperm to fertilize the female’s eggs. Sperm com-

mate and the benefit gained by mating with a preferred petition occurs when the ejaculates of two or more males

partner.[6] Costs include time and energy taken to choose compete for the fertilization of a given set of ova. In con-

the right males. Benefits include having offspring that trast cryptic female choice occurs when females them-

are more fit to survive and produce their own offspring, selves bias sperm use in favor of particular males.[12] Not

and having a mate that could potentially protect these only do the females control which sperm is used, but also

offspring better than others.[7] Mate choice is relatively the amount of sperm used. Females use both sperm dis-

costly to female marine iguanas as they spend around placement in which sperm is removed from storage by





1

From Wikipedia, the free encyclopedia Mate Choice





the incoming ejaculate of a copulating male, and sperm dominance, and the ability to pass along those traits to

incapacitation where incoming seminal fluids interfere offspring. Size assortive mating is one of the most com-

with resident sperm. Sperm dumping is the term used mon mating patterns in nature because of this.[15] This

to describe what the females will eventually do to used occurs in a wide range of animals from flatworms to

sperm or unused and unwanted sperm. In the spider: mammals. Size assortive mating may also result from fe-

opopae fosuma, there are several specialized muscles for males trying to match size. Prudent mate choice evolves

this function. The sperm storage site and the sperm fer- when small mates choosing larger mates would impose

tilization sites are separated from one another, allowing significant costs that outweigh the benefits of having a

females to eject sperm with the same genital mechanisms larger mate.[16] Specific size choice in sexual selection

that move stored sperm from the storage site to the fer- can lead to sexual dimorphism when the choice is only

tilization site.[13] This is primarily a function only exhib- made for one sex.[17] Male mate choice is also seen when

ited by spiders and also insects. dealing with specific size choice. When presented with

large and small potential mates, male pacific blue-eyed

Male Mate Choice/Sex Role fish spent more time and effort with the larger fe-

males.[18]

Reversal

There are certain situations in which the male is the one Sound

making the mate choice. In Polygynous societies, males Sound is another factor that can determine mate choice.

mate with multiple females. Males put forth little energy Sound is used not only to choose mates within a specific

towards mating most of the time because sperm produc- species, but is often used as a tool for species recognition

tion is easily done and large in quantity. On the other as well. In North American Tree Frogs, males form aggre-

hand females usually put a lot of energy towards mating gations and each male vocalizes from a relatively fixed

since eggs take much more energy and are a lot less ex- position during the calling period on a given night. The

pendable than sperm, which is constantly produced by female then usually moves close to or touches a calling

the body during a lifetime. Females usually end up taking male. The calls of different species are different each in

care of the young, also causing an energy imbalance be- their own ways, for example H. cinera produce unpulsed

tween the two sexes. This leaves the male better able to advertisement calls while H. versicolor readily increases

transfer genetic material between multiple females in a call duration by providing static impulses during the call.

short amount of time, allowing him to pass his genes to a This suggests that these calls are used at least partially

greater amount of offspring. Other times the number of for species recognition.[19] Females also preferred calls

females are high and the males are limited. In this case as that were at slightly different frequencies than the nor-

seen in the three spine stickleback, males choose the fe- mal frequency, around one to two kHz different. Male

males because they are the ones who have more of a se- field crickets use two acoustic signals in the context of

lection.[14] mating, one to attract females from a distance and a

courtship song when the female comes closer and makes



Evolutionary Conse- physical contact with the male.[20] Because of this mate

choice can occur at several different points. Females gen-

erally prefer the same type of frequency, and when the

quences frequency was changed on them artificially, they lost all

interest in the male. This signifies the importance of the

There are many different genetic factors that determine song on the attractiveness of the males in this species.

whether or not a male is a favorable mate to choose, or Birds are another type of animal in which songs are used

unfavorable. Actual genes cannot be seen when choos- for mate choice. In birds, males with a larger repertoire

ing mates, but there are many cues throughout the an- of songs are usually preferred. The Nutritional Strength

imal kingdom that help organisms predict the fitness of Hypothesis states that the amount of songs the bird

other organisms, making choosing mates more efficient knows is a good indicator of the bird’s quality because the

and more of a knowledgeable decision. Several factors development of the brain regions underlying song learn-

used by a wide variety of animals include: Size of mate, ing and production occurs when young birds typically

Sounds, Ornaments, Color, and dancing techniques. face nutritional and other stresses during post hatch de-

velopment. Therefore song repertoire length reflects

Size how well the bird fared during the post hatch period, and

might reflect how his offspring will fare during the post

Mates will usually choose organisms that are of larger hatch period.[21] Results show that even a relatively brief

size,

size excluding those who are smaller for a variety of period of undernutrition experienced early in life can af-

reasons. Some of these reasons include protection, more fect the outcome of song learning.



2

From Wikipedia, the free encyclopedia Mate Choice





Ornaments human mate choice that have no significance in the an-

imal kingdom. Money is one influence to choose a mate,

Another important factor in mate choice is ornaments and in our society it can play a rather large part. More

and their color. Ornaments come in various different wealthy individuals are usually seen as more able to pro-

forms, sizes, shapes, and colors. Depending upon the vide for one’s offspring. There is no such thing as physi-

species, the characteristics of the ornaments can be a tell cal money in the animal kingdom. Humans can also judge

tale sign towards compatibility and favorability. Resis- and measure expressed preferences before we are forced

tance to disease is a very important gene to pass down to make any choices via newspaper ads or online dating

to offspring and sometimes this characteristic can be dis- services.[28]

played by an ornament. During the breeding season, male

red jungle fowls with large combs, a sexually preferred

trait, have lower levels of lymphocytes, but greater cell-

mediated immunity indicated by a cutaneous hypersen-

See Also

sitivity response. Before the breeding season, both lym- • Sexual selection

phocyte proportion and cell-mediated immunity are pos- • Sexual conflict

itively associated with comb length.[22] A female in mak- • Polygyny

ing a mate choice obviously wants their offspring to have • Monogyny

better immunity against pathogens, so therefore will • Cryptic Choice

pick a mate who displays these characteristics over one

that does not. Organisms often do not have only one or-

nament to display. The California Quail has: a crest, cap

References

patch, white band, blue breast, black patch, tan patch, [1] Alcock, Jonathon (2009). Animal Behavior, Ninth

and a brown patch. Females choose their mates in this in- edition. Integrative and Comparative Biology.

stance from a combination of different features that dif- ISBN 10.1093/icb/icp058.

fer based upon their personal preference.[23] [2] Kokko, H; Brooks, Jennions, Morley (2003). "The

evolution of mate choice and mating biases".

Proceedings of the Royal Society of London. Series B:

Mate Choice Copying Biological Sciences 270 (1515): 653–664. doi:10.1098/

rspb.2002.2235. PMC 1691281. PMID 12769467.

Mate choice copying[24] is when one organism observes http://www.pubmedcentral.nih.gov/

and copies the mate choice of others. This occurs when articlerender.fcgi?tool=pmcentrez&artid=1691281.

mate choice is costly. This will increase the probability [3] Forsgren, E (1997). "Female sand gobies prefer good

of a mate being picked since the organism is copying fathers over dominant males". Proceedings of the

something that a successful mate performed. This allows Royal Society 264 (1368): 1283–1286. doi:10.1098/

a trait to spread non genetically. Male deep-snouted rspb.1997.0177.

pipefish exhibit this characteristic to enhance their own [4] Wong, B; Jennions (2003). "Costs influence male

fitness.[25] Females did not choose based upon observa- mate choice in a freshwater fish". Proceedings of the

tions of other females in this study. There is a higher rate Royal Society of London 270 (1): S36-S38. doi:10.1098/

of copying characteristics, than original characteristics rsbl.2003.0003.

being present. Females could not tell the difference be- [5] Wong, B; Jennions (2003). "Costs influence male

tween the two. mate choice in a freshwater fish". Proceedings of the

Royal Society of London 270 (1): S36-S38. doi:10.1098/

rsbl.2003.0003.

Human Mate Choice [6] Vitousek, M; Mitchell, Woakes, Niemack, Wikelski

(2007). "High costs of female choice in a lekking

Human beings also exhibit mate choice. There is substan- lizard". PloS one 2 (6). doi:10.1371/

tial evidence that in human mate choice, females direct- journal.pone.0000567.

ly select males based on male display of both physical [7] Vitousek, M; Mitchell, Woakes, Niemack, Wikelski

and behavioral traits.[26] In humans, the main form of (2007). "High costs of female choice in a lekking

attractiveness is the face, often the first thing that we lizard". PloS one 2 (6). doi:10.1371/

look at on another person and one of the only truly ex- journal.pone.0000567.

posed parts of the body. The parasite theory of sexual se- [8] Gwinner, H; Schwabl (2005). "Evidence for sexy

lection, states that men would prefer average and sym- sons in European starlings (Sturnus vulgaris)".

metrical faces in women, and that women would prefer Behavioral Ecology and Sociobiology 58 (4): 375–382.

largeness of the secondary sexual traits of the male’s doi:10.1007/s00265-005-0948-0.

face.[27] There are several other factors that influence



3

From Wikipedia, the free encyclopedia Mate Choice





[9] Garcia-Gonzalez, F; Simmons. "Good Genes and [20] Rebar, D; Bailey, Zuk (2009). "Courtship songʼs role

Sexual Selection in Dung Beetles (Onthophagus during female mate choice in the field cricket

taurus): Genetic Variance in Egg-to-Adult and Teleogryllus oceanicus". Behavioral Ecology 20 (6):

Adult Viability". PLoS ONE 6 (1). doi:10.1371/ 1307–1314.

journal.pone.0016233. [21] Nowicki (2002). "Brain development, song learning

[10] Garcia-Gonzalez, F; Simmons. "Good Genes and and mate choice in birds: a review and

Sexual Selection in Dung Beetles (Onthophagus experimental test of the "nutritional stress

taurus): Genetic Variance in Egg-to-Adult and hypothesis"". Journal of Comparative Physiology A:

Adult Viability". PLoS ONE 6 (1). doi:10.1371/ Neuroethology, Sensory, Neural, and Behavioral

journal.pone.0016233. Physiology 188 (11): 1003–1014. doi:10.1007/

[11] Edvardsson, M; Göran (2000). "Copulatory s00359-002-0361-3.

courtship and cryptic female choice in red flour [22] Zuk, M; Johnsen (1998). "Seasonal changes in the

beetles Tribolium castaneum". Proceedings of the relationship between ornamentation and immune

Royal Society of London 267 (1443): 559–563. response in red jungle fowl.". Proceedings of the

doi:10.1098/rspb.2000.1037. Royal Society of London 265 (1406): 1631–1635.

[12] Pilastro, A; Simonato, Bisazza, Evans (2004). doi:10.1098/rspb.1998.0481.

"CRYPTIC FEMALE PREFERENCE FOR COLORFUL [23] Calkins, D; Burley (2003). "Mate choice for multiple

MALES IN GUPPIES". Evolution 58 (3): 665–669. ornaments in the California quail, Callipepla

doi:10.1111/j.0014-3820.2004.tb01690.x. californica". Animal Behaviour. doi:10.1006/

[13] Burger, M.; Nentwig, Kropf (2003). "Complex anbe.2002.2041.

genital structures indicate cryptic female choice in [24] Widemo, M (2005). "Male but not female pipefish

a haplogyne spider (arachnida, araneae, oonopidae, copy mate choice". Behavioral Ecology 17 (2):

gamasomorphinae)". Journal of Morphology 1 (255): 255–259. doi:10.1093/beheco/arj021.

80–93. doi:10.1002/jmor.10048. [25] Widemo, M (2005). "Male but not female pipefish

[14] Sargent, R; Berghe (1986). Male Mate Choice in Fishes. copy mate choice". Behavioral Ecology 17 (2):

[15] Taborsky, B; GUYER, Taborsky (2009). "Size- 255–259. doi:10.1093/beheco/arj021.

assortative mating in the absence of mate choice". [26] Waynforth, D (2007). "Mate Choice Copying in

Animal Behaviour 77 (2): 439–448. Humans". Human Nature 18 (3). doi:10.1007/

[16] Taborsky, B; GUYER, Taborsky (2009). "Size- s12110-007-9004-2.

assortative mating in the absence of mate choice". [27] Grammer, K; Thornhill (1994)). "Human (Homo

Animal Behaviour 77 (2): 439–448. sapiens) facial attractiveness and sexual selection:

[17] Howard, R.D.; Martens, Innis, Drnevich, Hale the role of symmetry and averageness". Journal of

(1998)). "Mate choice and mate competition Comparative Psychology 108 (3): 233–242.

influence male body size in Japanese medaka". PMID 7924253.

Animal Behaviour 55 (5): 1151–1163. [28] Pawłwski, B; Dunbar (1999). "Impact of market

http://bilbo.bio.purdue.edu/~www_eco/faculty/ value on human mate choice decisions". Proceedings

howard/pdfs/medakaMS.pdf.. of the Royal Society of London. Series B: Biological

[18] Wong, B; Jennions (2004). "Sequential male mate Sciences 266 (1416): 281–285. doi:10.1098/

choice in a fish, the Pacific blue-eye rspb.1999.0634.

Pseudomugil signifer
and Sociobiology 56 (3): 253–256. doi:10.1007/

s00265-004-0775-8.

External links

[19] Gerhardt, H. C (1991)). "Female mate choice in • http://science.jrank.org/pages/48592/An-

treefrogs: static and dynamic criteria.". Animal Overview.html

Behaviour..





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