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Upwelling

Ecosystems

(Écosystèmes d’upwelling)

Structure and functioning of upwelling

ecosystems, under global change









ECO-UP

IRD Members

Name Theme Main ecosystem

Bertrand Arnaud Ecologist Humboldt

Cury Philippe Ecologist All

Demarcq Herve Biologist Benguela, Canary

Drapeau Laurent Computer scientist Benguela, Humboldt

Fréon Pierre Biologist All

Gerlotto Francois Biologist Humboldt

Lett Christophe Mathematical modelling Benguela, all

Machu Eric Oceanographer Canary

Mullon Christian Mathematical modelling All

Penven Pierrick Oceanographer Benguela, Humboldt

Roy Claude Oceanographer Benguela, Canary

Shin Yunne Ecologist Benguela

Students



Name Supervisor Period Ecosystem

Bertrand Sophie Gerlotto 2003-2005 Humboldt

Chang Nicolette Penven 2003-2006 Benguela

Kone Vamara Machu - Fréon 2003-2006 Benguela

Miller David Moloney - Lett 2003-2006 Benguela

Estrade Philippe Roy 2004-2007 Benguela

Mirabet Vincent Lett 2004-2007 General

Yemane Dawitt Shin - Field 2004-2007 Benguela

Neira Sergio Shannon - Cury 2005-2008 Benguela Humboldt

Sow Bamol Roy 2005-2008 Canary

Travers Morgane Shin - Field 2006-2008 Benguela

Brochier Timothee Lett - Fréon 2006-2008 Humboldt

Vergnon Remi Shin 2006-2008 General

Nieto Karen Demarcq 2006-2008 Camary-Humboldt

Main partnership Saint Quentin

UVSQ

Mullon









Brest Sete

IRD-Ifremer CRHMT

Roy Fréon, Cury,

Humboldt Canary Benguela Shin, Demarcq

IMARPE INRH UCT

Bertrand, Gerlotto Machu MCM

Drapeau, Lett, Penven

From IDYLE and ACTIVE to ECO-UP

IDYLE

Question: How adaptive strategies of pelagic

population do depend on the physical and

biological dynamics of upwelling systems ?

Field: Benguela ECOSYSTEMES D’UPWELLING

Approach:Modelling, GIS

Themes:Hydrodynamic modelling, IBM, GIS, remote Question: What are the effects of

sensing, retrospective analysis global changes on worldwide

Effectif: 5 scientists upwelling ecosystems?

Locations:Le Cap, Sète, Brest Field :Humboldt, Benguela, Canary

ACTIVE Approach:Comparative, Intégrated

Question: What is the effect of the behaviour of Themes:Climate et hydrodynamics,

pelagic fish about their catcahability ? productivity, hydrodynamics and

Field :Humboldt ecology, trophic structures, eco-

Approach: Acoustic surveys, modelling, data ethology of pelagic systems,

analysis fisheries economics

Themes: aggregative behaviour (tunas, FAD, Staff:13 scientists

schools and clusters of small pelagics), spatial Locations: Sète, Le Cap, Lima,

structuring of pelagic habitat (tuna) Casablanca, Brest, Saint Quentin

IRD Staff: 9 scientists

Locations:Valparaiso, La Réunion, Seychelles, Sète

Structure of the project

Field studies : Partnership

– Benguela EUR-OCEANS

– Canaries GLOBEC-SPACC

– Humboldt BCLME

PNEC

Scientific themes: BENEFIT

– Climate, environment, ATI-HUMBOLDT

hydrodynamics and transport

– production,

Teaching

– fish populations,

– fisheries Doctoral schools (UCT, Montpellier 2),

UBO Brest (Sciences de la mer),

VAS-ENSAR, Concepcion Chili,

Université de Dakar)

Summer schools

11 PhD students

Comparative approach

Canaries Benguela Humboldt (Californie)

Comparative approach



Subjets Scales



Reproductive strategies Spatial extension (the whole system)

Regime shifts (1500km x 300km)

Spatial dynamics Spatial grid : From méso-scale to sub-

Productivity turn-over meso-scale (~2 à 20 km)

Relationships between topography

and dynamics Time extension : Decennial (regime

Importance of large predators and shifts)

fishing Time step : form 2 days to year

Climate and physics



25 km

5 km

1 km



Hydrodynamics

ROMS

Physics and ecology

Transport

Lagrangian approaches









Bakun’s triad

Biological production

NPZD





Chlorophyll



a) NPZD









b) SeaWIFS

Fish communities



Trophic

structure





EwE

OSMOSE

Viability

approach

Fish communities



Trophic structure

Illustration of our approach



Question : Regime shifts

Hypothesis : Nature of

ecological controls (Top

Down, Bottom Up, Wasp-

Waist)

A tool : Ecosystem modelling

An expected result : Indicators

of ecosystem functioning

Fish communities



Trophic structure



GIS





Hake only

Sardine only

Hake & Sardine

Fish communities

Eco-Ethology



Spatial behaviour:



– of pelagic

populations (schools,

clusters)

– of plankton (patches,

3d)

– of fleets



as indicators of

ecosystems regimes

Fisheries









Towards an integrated model of the worldwide

system of small pelagic fisheries

Canary



Responsible : Eric Machu



Partners :

INRH (Maroc), LPA (Sénégal), JRC (Italie),

Université d’Alveiro (Portugal), LPO

(France), LEMAR (France), US

ACAPPELLA (France), US 025 (France)



Specific questions

Latitudinal gradient

Importance of the shelf

Ecosystem shifts

Fisheries

Iron and storms

Humboldt





Responsible : A. Bertrand



Partners:

IMARPE, University of San Marcos (Peru);

IFOP, University of Concepcion, INPESCA

(Chili);



Questions

ENSO

Productivity

Relationship between primary production and

pelagic production

Benguela







Responsible : L. Shannon, L. Drapeau



Partners : M&CM (South Africa), University of

Cape Town (South Africa), BCLME

(Angola, Namibia, South Africa)



Questions

Connexion Indian Ocean / Atlantic Ocean

Distance between spawning and nursery

areas

Differences North/South

Expected results



Methodology: towards the Ecoscope of upwelling

ecosystems



Generic tools for LME



Indicators for EAF



Atlas on line of upwelling ecosystems

Funding

IRD

13 full time positions

130.000 euros/year

3 PhD/year

French scientific programs

PNEC (16.000 euros/year)

ANR (30.000 euros/year)

European scientific programs

Euroceans (3 PhD/year)

Schedule

2004, From January to June : Submitting

the project

2004, November : Cape Town meeting,

focus on scientific questions

2005-2008 : Carrying out the project:

implementation of different approaches

on the 3 ecosystems; implementation of

the Atlas on line

Time table

Ecosys- 2005 2006 2007 2008

tem

Canary Partnership (1), Partnership (2) ROMS, NPZD, Synthesis

ROMS, EwE ROMS, Lagrangian, Lagrangian

IBM, NPZD, Vability IBM, OSMOSE,

GIS

Hu mboldt Accoustic Accoustic surveys, ROMS, NPZD, Synthesis

surveys. GIS, Eco -éthology EAF, GIS

ROMS, EwE surveys, Lagra n-

gian, NPZD,

OSMOSE, Viabi lity

Bengu ela ROMS, NPZD, ROMS, BCLME, L a- ROMS, BCLME Synthesis

IBM, Lagra n- grangian, IBM, Eco -

gian éthology surveys,

OSMOSE, EwE, Viability, GIS

GIS

All Watching sy s- Watching system, Climate change Synthesis

tem Sp atial indicators, scenarios,

Time series, EwE, Time series, Comparative

Bio -économy., Bio -eco, Book (2) analysis, Book

Book (1) (2)

— To be done — Done

Keywords

Comparative approach

Integrated approach



Structure and functionning

Change

APPENDICES

Cape Town – November 2004

Method



Climate and hydrodynamics: large scale Structure/Functioning

Climate and hydrodynamics: meso-scale Change

Production: bottom up Operational Oceanography

Trophodynamics: Trophic structure Comparison

Eco-ethology

Application to EAF, implications for

operational oceanography

Classified questions (1)



Climate and hydrodynamics: large scale

Are there regimes in the circulation and atmospheric dynamics of the upwelling systems? Are

there global climate features that explain the synchrony between the pelagic stocks

throughout the world?

What are the regime shifts in the Benguela (spatial) and Humboldt (temporal) in climate change?

What are the triggers? What maintain them?

What is the regional impact of climate change in upwelling systems, e.g. on its structure?

Climate and hydrodynamics: meso-scale

What are the determinants of meso-scale activities in upwelling systems? Is it the position of the

upwelling front? What are the forcing factors?

What is the importance of mesoscale structures (e.g. filaments, eddies, cool ridges, upwelling

front etc)? How to characterize them, spatially and dynamically? What is their variability?

Production: bottom up

What is the magnitude of interannual variability of primary production? What causes it? How to

relate intra- and interannual variability? What is the temporal scale of events: blooms,

collapse?

Which nutrient(s) limits the production in upwelling systems? Are upwelling systems food limited

or nutrient limited, is the transfer efficiency similar? For example, why the Humboldt system

is thought to be food limited?

Classified questions (2)





Trophodynamics: Trophic structure

How can we incorporate ecosystem considerations into fishing sectors management? How can

we quantify the effect of human activity on structure and functioning of the ecosystem?

How to reconcile single species and multispecies management approaches?

What is the role of fishing in driving regime shifts?

What is the role of trophic interactions in the success of recruitment?

Does the length of their food chains characterize the different ecosystems?

How can we characterize the role of a species in a foodweb?

How do we account for the high/low pelagic fish biomass in the various upwelling systems e.g.

southern/northern Benguela?

How to characterize the types of controls, and the switch between them? Do the trophic controls

depend on the trophic level?

What are the implications of spatial distributions (horizontal and vertical) in trophic interactions?

Classified questions (3)



Eco-ethology

How gregarious fish manage the multi-scale environmental variability intrinsic of upwelling

ecosystems

How collective fish structures can be indicators of fish interactions with their environment and

ecosystem status; this requires answer to a more specific question: what are the behavioural

mechanisms allowing the organisation of collective structures

What are the determinisms of fish cluster size, number and dynamics: population versus

environmental factors (eddies, etc.)

What are the interactions between predators (natural or fishers) and pelagic fish in terms of

spatial organisation

How can we integrate eco-ethological process in the adaptative management of resources in an

EAF framework

How the energetic inputs in a hierarchical system are transmitted across multi-scale physical and

biological organisation levels and shape fish and predators (fishers included) spatial

organisation

How can we characterise the spatial and dynamic scale-invariant patterns in hierarchical

structured upwelling ecosystems

Classified questions (4)





Application to EAF, implications for operational oceanography

How can operational oceanography contribute to EAF? (Feedback between what operational

oceanography can provide and what EAF requires and can use).

How can we build indicators that are amenable to management?

What are the consequences of regime shifts in fisheries management?

What are the relevant real-time indicators of changes and variability?

What indicators are requested for the management of an unstable resource? Which indicators

are relevant in that context?



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